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The hippocampal synaptic circuit is important for declarative memory. Information arrives in the hippocampus from entorhinal cortex through the perforant pathways, which provide both direct and indirect input to CA1 pyramidal neurons, the major output neurons of the hippocampus. (Arrows denote the direction of impulse flow.) In the indirect trisynaptic pathway neurons in layer II of entorhinal cortex send their axons through the perforant path to make excitatory synapses onto the granule cells of the dentate gyrus. The granule cells project through the mossy fiber pathway and make excitatory synapses with the pyramidal cells in area CA3 of the hippocampus. The CA3 cells excite the pyramidal cells in CA1 by means of the Schaffer collateral pathway. In the direct pathway neurons in layer III of entorhinal cortex project through the perforant path to make excitatory synapses on the distal dendrites of CA1 Source: Prefrontal Cortex, Hippocampus, and the Biology of Explicit Memory Storage, Principles of Neural Science, Fifth Editon pyramidal neurons without intervening synapses. Citation: Kandel ER, Schwartz JH, Jessell TM, Siegelbaum SA, Hudspeth AJ, Mack S. Principles of Neural Science, Fifth Editon; 2012 Available at: http://mhmedical.com/ Accessed: May 13, 2017 Copyright © 2017 McGraw-Hill Education. All rights reserved