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Animal Physiology Topic: Nervous System Badiozaman Sulaiman BSc.(UKM), MSc.(USM) – Molecular Biology Department of Zoology Faculty of Resource Science & Technology Universiti Malaysia Sarawak This OpenCourseWare@UNIMAS and its related course materials are licensed under a Crea<ve Commons A>ribu<on-‐NonCommercial-‐ShareAlike 4.0 Interna<onal License. At the end of the unit, you should be able to: 1. Discuss the coordina8on process in nervous system. 2. Compare the invertebrate and vertebrate nervous organiza8ons. 3. Describe the soma8c and autonomic nervous systems. 4. Explain the func8on, structure and types of neurons Nervous System • Consists of neural network that coordinate voluntary and involuntary ac<ons in the body. • Coordina<on is carried out via three overlapping func<ons; 1. Sensory input. 2. Integra8on. 3. Motor output. Coordina8on pathway Nervous organiza8on -‐ Invertebrate • The complexity of nervous organiza<on in an organism determines its capacity for integra<on. • Brain, which is the center for integra<on, is formed from concentra<on of neurons in the head region = cephaliza8on. • Simple invertebrates such as hydra (cnidarian) and starfish (echinoderm) are lacking of cephaliza<on. Cephaliza8on • Cephaliza<on start to appears in flatworm (planarian) and higher invertebrates (annelid, arthropod, mollusk). Nervous organiza8on -‐ Vertebrate • The vertebrate exhibits high degree of cephaliza<on and elaborate nervous system. • Organiza<on of the vertebrate nervous system can be classified into two components: 1. Central nervous system (CNS). 2. Peripheral nervous system (PNS). Nervous organiza8on – Vertebrate (con8nue) • The CNS consist of brain and spinal cord. – It receives sensory forma<on from the PNS, performs integra<on and ini<ates motor command. • The PNS consist of all the nerves that branched out from the CNS. – It carries sensory informa<on to the CNS, and sends motor command to the muscles and glands. CNS and PNS Control system • In the PNS, sensory informa<on is conveyed through the sensory (afferent) neurons and motor command is conveyed through the motor (efferent) neurons. • The motor neurons are func<onally divided into two categories: – Soma8c nervous system that regulate voluntary ac8ons such as walking. – Autonomic nervous system that regulate involuntary ac8ons such as hormone secre<on and peristalsis. Division of the autonomic system • The autonomic nervous system have of two divisions that control opposite reac<ons: – Parasympathe8c Division • Promotes responses that are associated with a relaxed state. • E.g. causes the pupil to constrict and promotes diges<on. – Sympathe8c Division • Promote responses that are associated with an ac8ve state. • Crucial in the fight or flight situa<on. • E.g. inhibits diges<on process and accelerates heartbeat. Neuron • Neuron (or nerve cell) is the structural and func8onal unit of the nervous system. • Sensory informa<on and motor command are conveyed through a neural network in the form of nerve impulse. • The neuron cons<tute of three parts: 1. Dendrites – the por<on of neuron that receives nerve impulse from sensory receptor or other neurons. 2. Cell body -‐ the main body that contains nucleus and organelles. 3. Axon – the por<on of the neuron that conveys nerve impulse to another neuron or target cells. Neuron (con8nue) • In myelinated neurons, the axons are covered with insula<ng layer called the myelin sheath. • The myelin sheath is formed from membranes of <ghtly spiraled neuroglia -‐ cells that provide support and nourishment to the neuron. – In PNS, the Schwann cells perform this func<on. – The gap between myelin sheath is called the nodes of Ranvier or neurofibril nodes. • The neurons are bundled together into nerve fibers. – A single nerve network contain several bundles of nerve fibers wrapped together. Nerve fibers Types of neurons • Sensory (afferent) neurons carry nerve impulses from sensory receptors to the CNS. – Sensory receptors may be a part of sensory neuron itself (e.g. pain receptor) or a specialized cell that forms a synapse with a sensory neuron. • Interneurons convey nerve impulses between various parts of the CNS. • Motor (efferent) neurons deliver nerve impulses from CNS to muscles or glands. – Have many dendrites and a single axon. – Cause muscle to contract or glands to secrete. Further reading: Reece, Urry, Cain, Wasserman, Minorsky & Jackson. (2013). Campbell Biology (10th edi<on). Benjamin Cummings. Willmer, Stone & Johnston. (2005). Environmental Physiology of Animals (2nd edi<on). Blackwell Publishing.