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PowerPoint Lecture Outlines to accompany Hole’s Human Anatomy and Physiology Tenth Edition Shier w Butler w Lewis Chapter 11 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. 11-1 Chapter 11 Nervous System II Meninges • membranes surrounding CNS • protect CNS • three layers • dura mater – outer, tough • arachnoid mater weblike • pia mater – inner, delicate 11-2 Meninges of the Spinal Cord 11-3 The Spinal Cord & Spinal Nerves • Together with brain forms the CNS • Functions – spinal cord reflexes – integration (summation of inhibitory and excitatory) nerve impulses – highway for upward and downward travel of sensory and motor information Meninges • The meninges are three coverings that run continuously around the spinal cord and brain (Figures 13.1a, 14.4a). – The outermost layer is the dura mater. – The middle layer is the arachnoid. – The innermost meninx is the pia mater, a thin, transparent connective tissue layer that adheres to the surface of the spinal cord and brain • Denticulate ligaments are thickenings of the pia mater that suspend the spinal cord in the middle of its dural sheath. Structures Covering the Spinal Cord • Vertebrae • Epidural space filled with fat • Dura mater – dense irregular CT tube • Subdural space filled with interstitial fluid • Arachnoid = spider web of collagen fibers • Subarachnoid space = CSF • Pia mater – thin layer covers BV – denticulate ligs hold in place Ventricles • interconnected cavities • within cerebral hemispheres and brain stem • continuous with central canal of spinal cord • filled with cerebrospinal fluid (csf) • lateral ventricles • third ventricle • fourth ventricle • cerebral aqueduct 11-4 Cerebrospinal Fluid • secreted by choroid plexus • circulates in ventricles, central canal of spinal cord, and subarachnoid space • completely surrounds brain and spinal cord • clear liquid • nutritive and protective • helps maintain stable ion concentrations in CNS 11-5 Spinal Cord Structure • extends foramen magnum to 2nd lumbar vertebra 11-6 Cross Section of Spinal Cord 11-7 Reflex Arcs Reflexes – automatic, subconscious responses to stimuli 11-9 Tracts of the Spinal Cord • Ascending tracts conduct sensory impulses to the brain • Descending tracts conduct motor impulses from the brain to motor neurons reaching muscles and glands 11-13 Ascending Tracts • fasciculus cuneatus • lateral spinothalamic 11-14 Corticospinal Tract 11-15 Brain Functions Major Parts • interprets sensations • cerebrum • two cerebral hemispheres • determines perception • diencephalon • stores memory • brain stem • reasoning • cerebellum • makes decisions • coordinates muscular movements • regulates visceral activities • determines personality 11-16 Brain Development Three Major Vesicles 1. Forebrain 2. Midbrain 3. Hindbrain Forebrain (prosencephalon) • anterior portion (telencephalon) • cerebrum • basal ganglia • posterior portion (diencephalon) • thalamus • hypothalamus • posterior pituitary • pineal gland 11-17 Brain Development Midbrain (mesencephalon) • midbrain Hindbrain (rhombencephalon) • anterior portion (metencephalon) • cerebellum • pons • posterior portion (myelencephalon) • medulla oblongata 11-18 Structure of Cerebrum • corpus callosum • connects hemispheres • convolutions • bumps or gyri • sulci • grooves • longitudinal fissure • separates hemispheres • transverse fissure • separates cerebrum from cerebellum 11-19 Lobes of Cerebrum • Frontal • Parietal • Temporal • Occipital • Insula 11-20 Functions of Cerebrum • interpretation • initiating voluntary movements • storing memory • retrieving memory • reasoning • center for intelligence and personality 11-21 Functional Regions of Cerebral Cortex Cerebral Cortex – thin layer of gray matter that constitutes the outermost portion of cerebrum; contains 75% of all neurons in nervous system 11-22 Motor Areas • Primary Motor Areas • frontal lobes • control voluntary muscles • Broca’s Area • anterior to primary motor cortex • usually in one hemisphere • controls muscles needed for speech • Frontal Eye Field • above Broca’s area • controls voluntary movements of eyes and eyelids 11-23 Motor Areas 11-24 Sensory Areas • Cutaneous Sensory Area • parietal lobe • interprets sensations on skin • Visual Area • occipital lobe • interprets vision • Auditory Area • temporal lobe • interprets hearing 11-25 Sensory Areas 11-26 Association Areas • regions of cortex that are not primary motor or primary sensory areas • widespread throughout the cerebral cortex • analyze and interpret sensory experiences • provide memory, reasoning, verbalization, judgment, emotions 11-27 Association Areas Frontal Lobe Association Areas • concentrating • planning • problem solving • judging Temporal Lobe Association Areas • remember visual scenes • remember music • remember complex patterns Parietal Lobe Association Areas • understanding speech • using words to express thought Occipital Lobe Association Areas • combine visual images with other sensory experiences 11-28 Memory Short Term • working memory • closed circuit • circuit is stimulated over and over • when impulse flow stops, memory disappears Long Term • changes structure and function of neurons • enhanced synaptic transmission 11-30 Basal Nuclei • masses of gray matter • deep within cerebral hemispheres • caudate nucleus, putamen, globus pallidus • produce dopamine • control certain muscular activities 11-31 Diencephalon • between cerebral hemispheres and brainstem • surrounds third ventricle • thalamus • hypothalamus • optic tracts • optic chiasm • infundibulum • posterior pituitary • mammillary bodies • pineal gland 11-32 Diencephalon Thalamus • gateway for sensory impulses heading to cerebral cortex • receives all sensory impulses (except smell) • channels impulses to appropriate part of cerebral cortex for interpretation Hypothalamus • maintains homeostasis by regulating visceral activities • links nervous and endocrine systems 11-33 Limbic System Consists of • portions of frontal lobe • portions of temporal lobe • hypothalamus • thalamus • basal nuclei • other deep nuclei Functions • controls emotions • produces feelings • interpret sensory impulses 11-34 Brain Stem Three Parts 1. Midbrain 2. Pons 3. Medulla Oblongata 11-35 Midbrain • between diencephalon and pons • contains bundles of fibers that join lower parts of brainstem and spinal cord with higher part of brain • cerebral aqueduct • cerebral peduncles – bundles of nerve fibers • corpora quadrigemina – centers for visual and auditory reflexes 11-36 Pons • rounded bulge on underside of brainstem • between medulla oblongata and midbrain • helps regulate rate and depth of breathing • relays nerve impulses to and from medulla oblongata and cerebellum 11-37 Medulla Oblongata • enlarged continuation of spinal cord • conducts ascending and descending impulses between brain and spinal cord • contains cardiac, vasomotor, and respiratory control centers • contains various nonvital reflex control centers (coughing, sneezing, vomiting) 11-38 Reticular Formation • complex network of nerve fibers scattered throughout the brain stem • extends into the diencephalon • connects to centers of hypothalamus, basal nuclei, cerebellum, and cerebrum • filters incoming sensory information • arouses cerebral cortex into state of wakefulness 11-39 Cerebellum • inferior to occipital lobes • posterior to pons and medulla oblongata • two hemispheres • vermis connects hemispheres • cerebellar cortex – gray matter • arbor vitae – white matter • cerebellar peduncles – nerve fiber tracts • dentate nucleus – largest nucleus in cerebellum • integrates sensory information concerning position of body parts • coordinates skeletal muscle activity • maintains posture 11-41 Peripheral Nervous System • Cranial nerves arising from the brain • Somatic fibers connecting to the skin and skeletal muscles • Autonomic fibers connecting to viscera • Spinal nerves arising from the spinal cord • Somatic fibers connecting to the skin and skeletal muscles • Autonomic fibers connecting to viscera 11-42 Structure of a Peripheral Nerve 11-43 Nerve Fiber Classification • Sensory Nerves – conduct impulses into CNS • Motor Nerves – conduct impulses to muscles or glands • Mixed Nerves – contain both sensory nerve fibers and motor nerve fibers; most nerves General somatic efferent fibers • carry motor impulses from CNS to skeletal muscles General somatic afferent fibers • carry sensory impulses to CNS from skin and skeletal muscles General visceral efferent fibers • carry motor impulses away from CNS to smooth muscles and glands General visceral afferent fibers • carry sensory impulses to CNS from blood vessels and internal organs 11-44 Nerve Fiber Classification Special somatic efferent fibers • carry motor impulses from brain to muscles used in chewing, swallowing, speaking, and forming facial expressions Special visceral afferent fibers • carry sensory impulses to brain from olfactory and taste receptors Special somatic afferent fibers • carry sensory impulses to brain from receptors of sight, hearing, and equilibrium 11-45 Cranial Nerves 11-46 Cranial Nerves I and II Olfactory (I) • sensory • fibers transmit impulses associated with smell Optic (II) • sensory • fibers transmit impulses associated with vision 11-47 Cranial Nerves III and IV Oculomotor (III) • primarily motor • motor impulses to muscles that • raise eyelids • move the eyes • focus lens •adjust light entering eye Trochlear (IV) • primarily motor • motor impulses to muscles that move the eyes 11-48 Cranial Nerve V Trigeminal (V) • mixed • opthalmic division • sensory from surface of eyes, tear glands, scalp, forehead, and upper eyelids • maxillary division • sensory from upper teeth, upper gum, upper lip, palate, and skin of face • mandibular division • sensory from scalp, skin of jaw, lower teeth, lower gum, and lower lip • motor to muscles of mastication and muscles in floor of mouth 11-49 Cranial Nerves VI and VII Abducens (VI) • primarily motor • motor impulses to muscles that move the eyes Facial (VII) • mixed • sensory from taste receptors • motor to muscles of facial expression, tear glands, and salivary glands 11-50 Cranial Nerves VIII and IX Vestibulocochlear (VIII) • sensory • sensory from equilibrium receptors of ear • sensory from hearing receptors Glossopharyngeal (IX) • mixed • sensory from pharynx, tonsils, tongue, and carotid arteries • motor to salivary glands and muscles of pharynx 11-51 Cranial Nerve X Vagus (X) • mixed • somatic motor to muscles of speech and swallowing • autonomic motor to viscera of thorax and abdomen • sensory from pharynx, larynx, esophagus, and viscera of thorax and abdomen 11-52 Cranial Nerves XI and XII Accessory (XI) • primarily motor • motor to muscles of soft palate, pharynx, larynx, neck, and back Hypoglossal (XII) • primarily motor • motor to muscles of the tongue 11-53 Spinal Nerves • mixed nerves • 31 pairs • 8 cervical (C1 to C8) • 12 thoracic (T1 to T12) • 5 lumbar (L1 to L5) • 5 sacral (S1 to S5) • 1 coccygeal (Co) 11-54 Spinal Nerves Dorsal root • axons of sensory neurons in the dorsal root ganglion Dorsal root ganglion • cell bodies of sensory neurons Ventral root • axons of motor neurons whose cell bodies are in spinal cord Spinal nerve • union of ventral root and dorsal root 11-55 Dermatome • an area of skin that the sensory nerve fibers of a particular spinal nerve innervate 11-56 Cervical Plexus Nerve plexus – complex networks formed by anterior branches of spinal nerves; fibers of various spinal nerves are sorted and recombined Cervical Plexus • C1-C4 • lies deep in the neck • supply muscles and skin of the neck • contribute to phrenic nerve 11-57 Brachial Plexus • C5-T1 • lies deep within shoulders • musculocutaneous nerves • supply muscles of anterior arms and skin of forearms • ulnar nerves • supply muscles of forearms and hands • supply skin of hands • radial nerves • supply posterior muscles of arms and skin of forearms and hands • axillary nerves • supply muscles and skin of superior, lateral, and posterior arms 11-58 Lumbosacral Plexus • T12 – S5 • extend from lumbar region into pelvic cavity • obturator nerves • supply adductors of thighs • femoral nerves • supply muscles and skin of thighs and legs • sciatic nerves • supply muscles and skin of thighs, legs, and feet 11-59 Autonomic Nervous System • functions without conscious effort • controls visceral activities • regulates smooth muscle, cardiac muscle, and glands • efferent fibers typically lead to ganglia outside CNS Two Divisions • sympathetic – prepares body for fight or flight situations • parasympathetic – prepares body for resting and digesting activities 11-60 Autonomic Nerve Fibers • all are motor (efferent) • preganglionic fibers • axons of preganglionic neurons • neuron cell bodies in CNS • postganglionic fibers • axons of postganglionic neurons • neuron cell bodies in ganglia 11-61 Sympathetic Division • thoracolumbar divison – location of preganglionic neurons • preganglionic fibers leave spinal nerves through white rami and enter paravertebral ganglia • paraverterbral ganglia and fibers that connect them make up the sympathetic trunk 11-62 Sympathetic Division • postganglionic fibers extend from sympathetic ganglia to visceral organs • postganglionic fibers usually pass through gray rami and return to a spinal nerve before proceeding to an effector • preganglionic fibers to adrenal medulla do not synapse with postganglionic neurons 11-63 Sympathetic Division 11-64 Parasympathetic Division • craniosacral division – location of preganglionic neurons • preganglionic fibers of the head in III, VII, and IX • ganglia are near or within various organs • preganglionic fibers of thorax and abdomen in X • short postganlionic fibers 11-65 Parasympathetic Division 11-66 Autonomic Neurotransmitters Cholinergic Fibers • release acetylcholine • preganglionic sympathetic fibers • preganglionic parasympathetic fibers • postganglionic parasympathetic fibers Adrenergic Fibers • release norepinephrine • postganglionic sympathetic fibers 11-67 Actions of Autonomic Neurotransmitters • depend on receptor Cholinergic receptors • bind to acetlycholine • muscarinic • excitatory • nicotinic • excitatory Adrenergic Receptors • bind to norepinephrine • alpha • different responses on various effectors • beta • different responses on various effectors 11-68 Actions of Autonomic Insert figure 11.39 Neurotransmitters 11-69 Control of Autonomic Activity • Controlled largely by CNS • Medulla oblongata regulates cardiac, vasomotor and respiratory activities • Hypothalamus regulates visceral functions • Limbic system and cerebral cortex control emotional responses 11-70 PowerPoint Lecture Outlines to accompany Hole’s Human Anatomy and Physiology Eleventh Edition Shier w Butler w Lewis Chapter 12 Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display. Senses Sensory Receptors • specialized cells or multicellular structures that collect information from the environment • stimulate neurons to send impulses along sensory fibers to the brain Sensation • a feeling that occurs when brain becomes aware of sensory impulse Perception • a person’s view of the stimulus; the way the brain interprets the information SENSORY RECEPTION • Sensory receptors convert stimulus energy to action potentials – Sensory receptors • Are specialized cells or neurons that detect stimuli – Sensory transduction converts stimulus energy into receptor potentials • Which trigger action potentials that are transmitted to the brain Sugar molecule Taste pore Tongue Taste bud 1 Sugar molecule (stimulus) Membrane of sensory receptor cell 2 Sensory receptor cells Signal transduction pathway Ion channels Sensory receptor cell 3 Ion Sensory neuron Receptor potential 4 Neurotransmitter Sensory neuron mV Action potential No sugar Figure 29.2A 5 Action potentials Sugar present – Action potential frequency • Reflects stimulus strength Sugar receptor “Sugar” interneuron “Salt” interneuron Salt receptor Brain Sensory neurons Taste bud No sugar Figure 29.2B Taste bud No salt Increasing sweetness Increasing saltiness • Mechanoreceptors – Mechanoreceptors • Respond to mechanical energy such as touch, pressure, and sound “Hairs” of receptor cell Neurotransmitter at synapse More neurotransmitter Less neurotransmitter Sensory neuron Action potentials Action potentials 1 Receptor cell at rest Figure 29.3B 2 Fluid moving in one direction 3 Fluid moving in other direction – Repeated stimulus • May lead to adaptation, a decrease in sensitivity • Specialized sensory receptors detect five categories of stimuli – A section of human skin • Reveals many types of sensory receptors Light Heat touch Pain Cold Hair Light touch Epidermis Dermis Figure 29.3A Hair Nerve Connective movement tissue Strong pressure Receptor Types Chemoreceptors • respond to changes in chemical concentrations Pain receptors (Nociceptors) • respond to tissue damage Thermoreceptors • respond to changes in temperature Mechanoreceptors • respond to mechanical forces Photoreceptors • respond to light Sensory Impulses • stimulation of receptor causes local change in its receptor potential • a graded electrical current is generated that reflects intensity of stimulation • if receptor is part of a neuron, the membrane potential may generate an action potential • if receptor is not part of a neuron, the receptor potential must be transferred to a neuron to trigger an action potential • peripheral nerves transmit impulses to CNS where they are analyzed and interpreted in the brain Sensations Projection process in which the brain projects the sensation back to the apparent source it allows a person to pinpoint the region of stimulation Sensory Adaptation • ability to ignore unimportant stimuli • involves a decreased response to a particular stimulus from the receptors (peripheral adaptations) or along the CNS pathways leading to the cerebral cortex (central adaptation) • sensory impulses become less frequent and may cease • stronger stimulus is required to trigger impulses Touch and Pressure Senses Free nerve endings • common in epithelial tissues • simplest receptors • sense itching Meissner’s corpuscles • abundant in hairless portions of skin; lips • detect fine touch; distinguish between two points on the skin Pacinian corpuscles • common in deeper subcutaneous tissues, tendons, and ligaments • detect heavy pressure and vibrations Temperature Senses Warm receptors • sensitive to temperatures above 25oC (77o F) • unresponsive to temperature above 45oC (113oF) Cold receptors • sensitive to temperature between 10oC (50oF) and 20oC (68oF) Pain receptors • respond to temperatures below 10oC • respond to temperatures above 45oC Sense of Pain • free nerve endings • widely distributed • nervous tissue of brain lacks pain receptors • stimulated by tissue damage, chemical, mechanical forces, or extremes in temperature • adapt very little, if at all Visceral Pain • pain receptors are the only receptors in viscera whose stimulation produces sensations • pain receptors respond differently to stimulation • not well localized • may feel as if coming from some other part of the body • known as referred pain Referred Pain • may occur due to sensory impulses from two regions following a common nerve pathway to brain Pain Nerve Pathways Acute pain fibers • A-delta fibers •thin, myelinated • conduct impulses rapidly • associated with sharp pain • well localized Chronic pain fibers • C fibers •thin, unmyelinated • conduct impulses more slowly • associated with dull, aching pain • difficult to pinpoint Regulation of Pain Impulses Thalamus • allows person to be aware of pain Cerebral Cortex • judges intensity of pain • locates source of pain • produces emotional and motor responses to pain Pain Inhibiting Substances • enkephalins • serotonin • endorphins Proprioceptors • mechanoreceptors • send information to spinal cord and CNS about body position and length and tension of muscles • Main kinds of proprioreceptors • Pacinian corpuscles – in joints • muscle spindles – in skeletal muscles* • Golgi tendon organs – in tendons* *stretch receptors Stretch Receptors Sense of Smell Olfactory Receptors • chemoreceptors • respond to chemicals dissolved in liquids Olfactory Organs • contain olfactory receptors and supporting epithelial cells • cover parts of nasal cavity, superior nasal conchae, and a portion of the nasal septum Olfactory Receptors Olfactory Nerve Pathways Once olfactory receptors are stimulated, nerve impulses travel through • olfactory nerves olfactory bulbs olfactory tracts limbic system (for emotions) and olfactory cortex/temporal lobe (for interpretation) Olfactory Stimulation • olfactory organs located high in the nasal cavity above the usual pathway of inhaled air • olfactory receptors undergo sensory adaptation rapidly • sense of smell drops by 50% within a second after stimulation Sense of Taste Taste Buds • organs of taste • located on papillae of tongue, roof of mouth, linings of cheeks and walls of pharynx Taste Receptors • chemoreceptors • taste cells – modified epithelial cells that function as receptors • taste hairs –microvilli that protrude from taste cells; sensitive parts of taste cells Taste Receptors Taste Nerve Pathways Sensory impulses from taste receptors travel along • cranial nerves to • medulla oblongata to • thalamus to • gustatory cortex / parietal lobes (for interpretation) Hearing Ear – organ of hearing Three Sections • External • Middle • Inner External Ear • auricle • collects sounds waves • external auditory meatus • lined with ceruminous glands • carries sound to tympanic membrane • terminates with tympanic membrane • tympanic membrane • vibrates in response to sound waves Middle Ear • tympanic cavity • air-filled space in temporal bone • auditory ossicles • vibrate in response to tympanic membrane • malleus, incus, and stapes • oval window • opening in wall of tympanic cavity • stapes vibrates against it to move fluids in inner ear Auditory Tube • eustachian tube • connects middle ear to throat • helps maintain equal pressure on both sides of tympanic membrane • usually closed by valve-like flaps in throat Inner Ear • complex system of labyrinths • osseous labyrinth • bony canal in temporal bone • filled with perilymph • membranous labyrinth • tube within osseous labyrinth • filled with endolymph Inner Ear Three Parts of Labyrinths • cochlea • functions in hearing • semicircular canals • functions in equilibrium • vestibule • functions in equilibrium Cochlea Scala vestibuli • upper compartment • leads from oval window to apex of spiral • part of bony labyrinth Scala tympani • lower compartment • extends from apex of the cochlea to round window • part of bony labyrinth Cochlea Cochlear duct • portion of membranous labyrinth in cochlea Vestibular membrane • separates cochlear duct from scala vestibuli Basilar membrane • separates cochlear duct from scala tympani Organ of Corti • group of hearing receptor cells (hair cells) • on upper surface of basilar membrane • different frequencies of vibration move different parts of basilar membrane • particular sound frequencies cause hairs of receptor cells to bend • nerve impulse generated Auditory Nerve Pathways Summary of the Generation of Sensory Impulses from the Ear Equilibrium Static Equilibrium • vestibule • sense position of head when body is not moving Dynamic Equilibrium • semicircular canals • sense rotation and movement of head and body Vestibule • Utricle • communicates with saccule and membranous portion of semicircular canals • Saccule • communicates with cochlear duct • Macula • hair cells of utricle and saccule Macula • responds to changes in head position • bending of hairs results in generation of nerve impulse Semicircular Canals • three canals at right angles • ampulla • swelling of membranous labyrinth that communicates with the vestibule • crista ampullaris • sensory organ of ampulla • hair cells and supporting cells • rapid turns of head or body stimulate hair cells Crista Ampullaris Sight Visual Accessory Organs • eyelids • lacrimal apparatus • extrinsic eye muscles Eyelid • palpebra • composed of four layers • skin • muscle • connective tissue • conjunctiva • orbicularis oculi - closes • levator palperbrae superioris – opens • tarsal glands – secrete oil onto eyelashes • conjunctiva – mucous membrane; lines eyelid and covers portion of eyeball Lacrimal Apparatus • lacrimal gland • lateral to eye • secretes tears • canaliculi • collect tears • lacrimal sac • collects from canaliculi • nasolacrimal duct • collects from lacrimal sac • empties tears into nasal cavity Extrinsic Eye Muscles Superior rectus • rotates eye up and medially Inferior rectus • rotates eye down and medially Medial rectus • rotates eye medially Extrinsic Eye Muscles Lateral rectus • rotates eye laterally Superior oblique • rotates eye down and laterally Inferior oblique • rotates eye up and laterally Structure of the Eye • hollow • spherical • wall has 3 layers • outer fibrous tunic • middle vascular tunic • inner nervous tunic Outer Tunic Cornea • anterior portion • transparent • light transmission • light refraction Sclera • posterior portion • opaque • protection Middle Tunic Iris • anterior portion • pigmented • controls light intensity Ciliary body • anterior portion • pigmented • holds lens • moves lens for focusing Choroid coat • provides blood supply • pigments absorb extra light Anterior Portion of Eye • filled with aqueous humor Lens • transparent • biconvex • lies behind iris • largely composed of lens fibers • elastic • held in place by suspensory ligaments of ciliary body Ciliary Body • forms internal ring around front of eye • ciliary processes – radiating folds • ciliary muscles – contract and relax to move lens Accommodation • changing of lens shape to view objects • To focus, a lens changes position or shape – Focusing • Involves changing the shape of the lens Choroid Ciliary muscle contracted Ligaments slacken Retina Light from a near object (diverging rays) Near vision (accommodation) Ciliary muscle relaxed Ligaments pull on lens Light from a distant object (parallel rays) Figure 29.6 Distance vision Lens Iris • composed of connective tissue and smooth muscle • pupil is hole in iris • dim light stimulates radial muscles and pupil dilates • bright light stimulates circular muscles and pupil constricts Aqueous Humor • fluid in anterior cavity of eye • secreted by epithelium on inner surface of the ciliary body • provides nutrients • maintains shape of anterior portion of eye • leaves cavity through canal of Schlemm Inner Tunic • retina • contains visual receptors • continuous with optic nerve • ends just behind margin of the ciliary body • composed of several layers • macula lutea – yellowish spot in retina • fovea centralis – center of macula lutea; produces sharpest vision • optic disc – blind spot; contains no visual receptors • vitreous humor – thick gel that holds retina flat against choroid coat Posterior Cavity • contains vitreous humor – thick gel that holds retina flat against choroid coat Major Groups of Retinal Neurons • receptor cells, bipolar cells, and ganglion cells - provide pathway for impulses triggered by photoreceptors to reach the optic nerve • horizontal cells and amacrine cells – modify impulses Layers of the Eye Visual Receptors Rods • long, thin projections • contain light sensitive pigment called rhodopsin • hundred times more sensitive to light than cones • provide vision in dim light • produce colorless vision • produce outlines of objects Cones • short, blunt projections • contain light sensitive pigments called erythrolabe, chlorolabe, and cyanolabe • provide vision in bright light • produce sharp images • produce color vision Visual Pigments Rhodopsin • light-sensitive pigment in rods • decomposes in presence of light • triggers a complex series of reactions that initiate nerve impulses • impulses travel along optic nerve Pigments on Cones • each set contains different lightsensitive pigment • each set is sensitive to different wavelengths • color perceived depends on which sets of cones are stimulated • erythrolabe – responds to red • chlorolabe – responds to green • cyanolabe – responds to blue Visual Nerve Pathway