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Transcript
The Auditory Pathway
• This graphic depicts the events in the
stimulation of auditory receptors, from
channeling sound waves into the external ear
and onto the TM, to the transduction of those
vibrations
into local
receptor
potentials
Malleus Incus
Stapes vibrating Helicotrema
in oval window
Cochlea
Sound waves
Perilymph
3
7
4
5
1
2
6
9
External auditory
canal
8
Scala
tympani
Scala
vestibuli
Basilar
membrane
8
Spiral organ
(organ of Corti)
Tectorial membrane
Vestibular membrane
Cochlear duct
(contains endolymph)
Tympanic
membrane
Secondary tympanic
membrane vibrating
in round window
Middle ear
Auditory tube
The Auditory Pathway
• The cell bodies of the sensory neurons are
located in the spiral ganglia. Nerve impulses
pass along the axons of these neurons, which
form the cochlear
branch of the
vestibulocochlear (VIII)
nerve
The Auditory Pathway
• The nerve impulses follow CN VIII en route to the
medulla, pons, midbrain, and thalamus, and finally
to the primary auditory cortex in the temporal
lobe. Slight differences in the timing of nerve
impulses arriving from the
two ears at the superior
olivary nuclei in the
pons allow us to
locate the source
of a sound
Equilibrium
• Equilibrium is another function of the inner ear controlled by the vestibular apparatus (the
saccule and utricle of the vestibule, and the 3
semicircular canals)
– Static equilibrium refers to a
state of balance relative to
the force of gravity
– Dynamic equilibrium
involves the maintenance
of balance during sudden movements
Static Equilibrium
• Static equilibrium is controlled by the sensory
hairs within the macula of the utricle
and saccule
The Inner Ear
• Movements of the hair cells in contact with the tectorial
membrane transduce mechanical vibrations into
electrical signals which generate nerve impulses along
the cochlear branch of CN VIII
Static Equilibrium
• An otolithic membrane, studded with dense
calcium carbonate crystals (otoliths), responds
to gravity when head position is changed
– This movement opens
transduction channels
in the hair cells,
producing local potentials
which summate to
form nerve AP