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Ocular Dominance Columns QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Binocular Vision • LGN receive input from both eyes, but these are in separate layers. • Individual LGN neurons are monoocular driven. • Ocular dominance columns in layer IV of the visual cortex. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. How are neurons with different receptive fields arranged within the striate cortex? • A column of cells has similar response properties. • Adjacent columns have similar response properties. • Same orientation is repeated at approx. 1 mm horizontal intervals (orientation preference). QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Ocular Dominance Columns • The striate cortex is composed of repeating units that contain all the neuronal components to analyze a small region of visual space for a variety of different stimulus attributes. Ocular Dominance Columns • Neurons respond to a single eye at the extreme of a column, middle area responds to both eyes. Within the primary visual pathway. • LGN layers • Different cell size. • There are separate functional streams for conveying information from different types of retinal ganglion cells to initial stages of cortical processing Different cell size. • MAGNOCELLULAR STREAM: for detecting rapidly changing stimuli (ventral) • PARVOCELLULAR STREAM: high acuity and colour vision (dorsal) • KONIOCELLULAR STREAM: shares colour vision with parvocellular stream. (interlaminar) QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. LGN Magnocellular & Parvocellular Streams • M ganglion cells and P ganglion cells terminate in separate layers. • Synapse onto neurons projecting to separate strata of striate cortex layer IV. M ganglion cells • Large receptive fields • Axons have fast conduction velocity • Respond transiently to stimulus presentation. P ganglion cells • • • • • Small receptive fields Slower conduction velocity Sustained response Can transmit colour information. Center/surround, red/green; green/red Koniocellular Pathway • Evidence for input from short wavelength color. • LGN interlaminar areas