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The Cerebellum SHANDONG UNIVERSITY Liu Zhiyu Position: lies above and behind the medulla oblongata and pons; occupies posterior cranial fossa Cerebellum External features Three peduncles Inferior cerebellar peduncle connect with medulla and spinal cord, contain both afferent and efferent fibers External features Three peduncles Middle cerebellar peduncle connect with pons, contain afferent fibers Superior cerebellar peduncle connect with midbrain, contain mostly efferent fibers External features Tonsil of cerebellum two elevated masses on inferior surface of hemisphere, just nearby foramen magnum Lobes of cerebellum Two deep fissures Primary fissure Posterolateral fissure Three lobs 1. Flocculonodular lobe flocculus and nodule 2. Anterior lobe 3. Posterior lobe corpus of cerebellar Lobes Anterior lobe Primary fissure corpus of Cerebellum Posterior lobe Flocculus Posterolateral fissure nodule Flocculonodular lobe Ⅱ. Internal structures Gray matter Cerebellar cortex Cerebellar nuclei Dentate nucleus Fastigial nucleus Interposed nuclei Emboliform nucleus Globose nucleus Ⅱ. Internal structures White matter— medullary center Cerebellar cortex Fastigial nucleus Globose nucleus Dentate nucleus medullary center Emboliform nucleus Three functional divisions Spinocerebellum ( paleocerebellum ) : vermis and intermediate zone Vestibulocerebellum ( archicerebellum): Flocculonodular lobe and vestibular nuclei lateral zone intermediate zone Cerebrocerebellum (neocerebellum ): lateral zone vermis Flocculonodular lobe Ⅲ. Connections and peduncles of cerebellum Middle cerebellar peduncle is composed of fibres which arises in the opposite side of the pontine nuclei and pass to the erebellum, transmit impulses which reach the the pons from the cerebral cortex. Ⅲ. Connections and peduncles of cerebellum inferior cerebellar peduncle This peduncle consists mainly of afferent fibres to the cerebellum,inlude posterior spinocerebellar tract; the fibers from the inferior olivary nucleus, the fibers from vestibular nuclei and nerve. It also transmits efferent fibres to the medulla oblongata. Ⅲ. Connections and peduncles of cerebellum superior cerebellar peduncle This is the principal efferent pathway from the cerebellum, its fibres arises mainly from the dentate nucleus, and passes anterosuperiorly to the opposite thalamus and red nucleus of the midbrain. IV. Connections and function of cerebellum Vestibulocerebellum Afferents fibers receive input from vestibular nuclei and vestibular n.. Efferents fibers : to the vestibular nucleus → vestibulospinal tract → anterior horn.of spinal cord. → medial longitudinal fasciculus → motor nuclei asociated with movement of eyeball。 Function: involved in eyeball movements and maintain the balance of the body IV. Connections and function of cerebellum Spinocerebellum Afferent fibers: receive somatic sensory information via spinocerebellar tracts. Efferent fibers: to the fastigial nucleus→ vestibular nuclei → vestibulospinal tract → motor neurons of anterior horn IV. Connections and function of cerebellum Spinocerebellum Efferent fibers: to the interposed nuclei中间 ① contralateral red nucleus → rubrospinal tract →motor neurons of anterior horn of spinal cord ② contralateral thalamus→cerebral cortex IV. Connections and function of cerebellum Spinocerebellum Function: play an important role in control of muscle tone and coordination of muscle movement on the same side of the body IV. Connections and function of cerebellum Cerebrocerebellum Afferents: receives input from the cerebral cortex via the pontine nuclei middle cerebellar peduncle Efferents: to dentate nucleus → contralateral thalamus → primary motor cortex → corticospinal tract → motor neurons of anterior horn Function: coordination of muscle movement of limbs.