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NAME DATE PERIOD Other Genetic Crosses J. Levine and K. Miller; 2002 Not all genes show simple patterns of dominant and recessive alleles. In most organisms, genetics is more complicated, because the majority of genes have more than two alleles. In addition, many important traits are controlled by more than one gene. Some alleles are neither dominant nor recessive, and many traits are controlled by multiple alleles or multiple genes. Incomplete Dominance A cross between two four o’clock plants shows one of these complications. The F1 generation produced by a cross between red-flowered (R1R1) and white-flowered (R2R2) plants consists of pink-colored flowers (R1R2). Which allele is dominant in this case? Neither one. Cases in which one allele is not completely dominant over another are called incomplete dominance. In incomplete dominance, the heterozygous phenotype is somewhere in between the two homozygous phenotype. 2 R R2 R1 R1 R2 R1 R2 R1 R1 R2 R1 R2 R1 R1= Red Flowers R2 R2= White Flowers R1 R2 = Pink Flowers Sex-Linked Traits Hemophilia, coat color in cats, color blindness are some genes located on the X chromosome, thus referred to as Sex-Linked Traits. While males have an X and Y sex chromosome, females have two X’s. In cats, for example the color for their coat is located on the X chromosome. One X chromosome may have the allele for the color orange, while the other X may have the allele for black. So it’s an X-linked polygenic trait. Since females have two X chromosomes, they could haveboth of those colors. If you see a cat with 3 colors: white, black, and orange, it’s almost certainly a female. Calico cats that are male are rare and infertile (XXY). Polygenic Traits Many traits are produced by the interaction of several genes. Traits controlled by two or more genes are said to be polygenic traits, which means, “having many genes.” Polygenic traits often show a wide range of phenotypes. For example, the wide range of skin color in humans comes about partly because more than four different genes probably control this trait. What are these examples of? Codominance QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. A similar situation is codominance, in which both alleles contribute to the phenotype of the organism. For example, in cattle the allele for red hair (RR) is codominant with the allele for white hair (WW). Cattle with both alleles are called “roan,” or reddish brown and white (RW), because their coats are a mixture of both red and white hairs. Q ui ck Ti m e ™ an d a T IF F ( Un co m p re ss ed ) d ec om pr e ss or a re ne ed ed t o s ee th i s pi c tu r e. Qu i ck Ti m e ™ an d a TI F F ( Un co m pr e ss ed ) d ec om p r ess or ar e n ee de d t o s ee t h is pi ct u re . Qu i ck Ti m e ™ an d a TI F F ( Un co m pr e ss ed ) d ec om p r es so r ar e n ee de d t o s ee t h is pi ct u re . Multiple Alleles Many genes have more than two alleles and are therefore said to have multiple alleles. This does not mean that an individual can have more than two alleles. It only means that more than two possible exist in a population. One of the best-known examples is coat color in rabbits. A rabbit’s coat color is determined by a single gene that has at least four different alleles. The four known alleles display a pattern of simple dominance that can produce four possible coat colors. Many other genes have multiple alleles, including the human genes for blood type. 841167681 Standard 8.0 – RT, RI, Monitoring, Feedback, Mastery © «GreetingLine» High School