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Download THE PLANT BODY AND PLANT CELLS
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THE PLANT BODY AND PLANT CELLS Plant body overview Shoot system --generally aerial --stem: axis of shoot --apex; apical bud: growing point --cotyledons --attached leaves at nodes (note internodes) blade, petiole arrangement on stem variable (opposite, alternate, etc) axil (with axillary bud) --axillary buds: dormant or growing --branch: from axillary bud --flowers: apical or axillary --apex; apical bud: growing point --stem: axis of shoot -- leaves: attached at nodes internode blade petiole axil (with axillary bud) alternate arrangement opposite arrangement --cotyledons --flowers: apical or axillary --axillary buds: dormant or growing b r a n ch from axillary bud Root system --generally underground --highly branched (lateral roots, but no nodes) --2 types: fibrous and tap root systems --growth at apices of all branches Note the two main types of flowering plants: dicots and monocots Dicots Monocots Seed leaves, Leaf venation cotyledons Two Reticulate One Parallel Bougainvilleas are dicots. Secondary growth Flower part multiples Yes Unusual 4s and 5s 3s Orchids are monocots. The bean seed is a dicot seed: two cotyledons The rice seed is a monocot seed: one cotyledon Dicots have reticulate venation Monocots have parallel venation Types of plant cells (tissues) Parenchymal cell (parenchyma) --cell wall: cellulose fibrils, connected by shorter, more random carbohydrate chains (pectin, hemicellulose), strong, elastic --plasma membrane --vacuole, tonoplast --nucleus, nucleolus, chromatin, nuclear envelope --plastids: chloroplasts, chromoplasts, amyloplasts, proplastids --mitochondria --endoplasmic reticulum --Golgi, called dictyosomes --ribosomes, polysomes --"standard, non-specialized" plant cell; photosynthetic (in green shoots); for storage (in shoots and roots); grows, provides some structure (turgid, stiff) Osmosis and turgor Because the plasma membrane is differentially permeable, allowing water to cross more easily than solutes, water moves from the side with the lowest solute concentration to the side with the highest concentration. Osmosis and turgor: In a plant, turgor makes a parenchymal cells able to provide structure. Assume cells with p.m., cell wall, solutes in cytoplasm, external dilute solution Solute concentrations unbalanced Water concentrations (activities) unbalanced Water flows in along activity gradient (osmosis) Cell volume rises, membrane pushes against wall (turgor pressure) Wall resists stretching, pushes back elastically (wall pressure) Equilibrium exists when water flow due to osmosis = water flow due to wall pressure So pressure of wall on membrane keeps cell from lysing in pure water or dilute solution; but also, pressure of membrane on wall provides structure (like air in tire); also, pressure provides force for controlled growth (if wall "gives")] Epidermal cells (epidermis) --thin, broad cells that form skin over leaves, primary stems, roots --secretes cuticle on outer surface (very hydrophobic): cutin (long chain fatty acids bound to cell wall) and waxes (long-chain fatty acids esterified to long chain alcohols) CH3-CH2-CH2-CH2-----CO-O-CH2-CH2-CH2---CH3 --produces secondary compounds, stores them in vacuole: absorb UV light, resist insects, fungi, etc. Collenchymal cells (collenchyma) --extra primary cell walls --support (e.g. leaf petioles [celery]), sometimes have chloroplasts Fibers, schlereids (schlerenchyma) --secondary cell wall, containing cellulose + lignin (highly crosslinked polyphenolic compound, very hard, hydrophobic), in strands or sheets under primary cell wall --autolysis of cell contents when wall finished --wall provides support against compression and expansion (even in low water-potential environment); resistance to grazing (seed coat) --with larger lumen, can serve as water conduit Collenchyma cell Summary •Shoot system •Root system •Dicots vs monocots •Basic plant cells and tissues • Parenchyma • Collenchyma • Fibers and sclereids •Osmosis and turgor