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Essentials of Human Anatomy Special Senses Dr Fadel Naim Ass. Prof. Faculty of Medicine IUG Special Senses • sensory receptors are within large, complex sensory organs in the head • smell in olfactory organs • taste in taste buds • hearing and equilibrium in ears • sight in eyes Sense of Smell Olfactory Receptors • chemoreceptors • respond to chemicals dissolved in liquids Olfactory Organs • contain olfactory receptors and supporting epithelial cells • cover parts of nasal cavity, superior nasal conchae, and a portion of the nasal septum Olfactory Receptors Olfactory Nerve Pathways Once olfactory receptors are stimulated, nerve impulses travel through • olfactory nerves olfactory bulbs olfactory tracts limbic system (for emotions) and olfactory cortex (for interpretation) Olfactory Stimulation • olfactory organs located high in the nasal cavity above the usual pathway of inhaled air • olfactory receptors undergo sensory adaptation rapidly • sense of smell drops by 50% within a second after stimulation Sense of Taste Taste Buds • organs of taste • located on papillae of tongue, roof of mouth, linings of cheeks and walls of pharynx Taste Receptors • chemoreceptors • taste cells – modified epithelial cells that function as receptors • taste hairs –microvilli that protrude from taste cells; sensitive parts of taste cells Taste Receptors Taste Sensations Four Primary Taste Sensations • sweet – stimulated by carbohydrates • salty – stimulated by salts • sour – stimulated by acids • bitter – stimulated by many organic compounds Spicy foods activate pain receptors Taste Nerve Pathways Sensory impulses from taste receptors travel along • cranial nerves to • medulla oblongata to • thalamus to • gustatory cortex (for interpretation) Hearing Ear – organ of hearing Three Sections • External • Middle • Inner External Ear • auricle • collects sounds waves • external auditory meatus • lined with ceruminous glands • carries sound to tympanic membrane • terminates with tympanic membrane • tympanic membrane • vibrates in response to sound waves Middle Ear • tympanic cavity • air-filled space in temporal bone • auditory ossicles • vibrate in response to tympanic membrane • malleus, incus, and stapes • oval window • opening in wall of tympanic cavity • stapes vibrates against it to move fluids in inner ear Auditory Tube • eustachian tube • connects middle ear to throat • helps maintain equal pressure on both sides of tympanic membrane • usually closed by valve-like flaps in throat Inner Ear • complex system of labyrinths • osseous labyrinth • bony canal in temporal bone • filled with perilymph • membranous labyrinth • tube within osseous labyrinth • filled with endolymph Inner Ear Three Parts of Labyrinths • cochlea • functions in hearing • semicircular canals • functions in equilibrium • vestibule • functions in equilibrium Cochlea Scala vestibuli • upper compartment • leads from oval window to apex of spiral part of bony labyrinth Scala tympani • lower compartment • extends from apex of the cochlea to round window • part of bony labyrinth Cochlea Cochlear duct • portion of membranous labyrinth in cochlea Vestibular membrane • separates cochlear duct from scala vestibuli Basilar membrane • separates cochlear duct from scala tympani Organ of Corti • group of hearing receptor cells (hair cells) • on upper surface of basilar membrane • different frequencies of vibration move different parts of basilar membrane • particular sound frequencies cause hairs of receptor cells to bend • nerve impulse generated Auditory Pathway Cochlear branch of CN VIII To cochlear nucleus of medulla To inferior colliculus of opposite side of midbrain To thalamus To auditory cortex Vestibular Complex: • Semicircular canals with ampullae (mutually perpendicular) • Saccule and utricle (= fill up vestibule) Two Receptor Organs: Maculae of Vestibule (or: macula of saccule plus macula of utricle) Macula • responds to changes in head position • bending of hairs results in generation of nerve impulse Crista Ampullaris •sensory organ of ampulla • hair cells and supporting cells • rapid turns of head or body stimulate hair cells Equilibrium • Rotation of the head causes endolymph within the semicircular canal to push against the cupula covering the hair cells, resulting in bending of their stereocilia and the initiation of a nerve impulse. Equilibrium Static Equilibrium • vestibule • sense position of head when body is not moving Dynamic Equilibrium • semicircular canals • sense rotation and movement of head and body The Sense of Vision • Visual receptors (photoreceptors) in the eyes to detect light, color, and movement. • Accessory structures of the eye. – provide a superficial covering over its anterior exposed surface (conjunctiva) – prevent foreign objects from coming into contact with the eye (eyebrows, eyelashes, and eyelids) – keep the exposed surface moist, clean, and lubricated (lacrimal glands) Lacrimal Apparatus • lacrimal gland • lateral to eye • secretes tears • canaliculi • collect tears • lacrimal sac • collects from canaliculi • nasolacrimal duct • collects from lacrimal sac • empties tears into nasal cavity Structure of the Eye • hollow • spherical • wall has 3 layers • outer fibrous tunic • middle vascular tunic • inner nervous tunic Cavities and Chambers of the Eye • The internal space of the eye is subdivided by the lens into two separate cavities. – anterior cavity – posterior cavity Cavities and Chambers of the Eye • The anterior cavity is – the space anterior to the lens and posterior to the cornea • The iris of the eye subdivides the anterior cavity further into two chambers. – anterior chamber is between the iris and cornea – posterior chamber is between the lens and the iris Vitreous Humor • Posterior cavity is posterior to the lens and anterior to the retina. • Transparent, gelatinous vitreous body which completely fills the space between the lens and the retina. Optic Disc • Optic disc lacks photoreceptors. • Called the blind spot because no image forms there. • Just lateral to the optic disc is a rounded, yellowish region of the retina called the macula lutea containing a pit called the fovea centralis (the area of sharpest vision). – contains the highest proportion of cones and almost no rods Visual Pathways • Each optic nerve conducts visual stimulus information. • At the optic chiasm, some axons from the optic nerve decussate. • The optic tract on each side then contains axons from both eyes. • Visual stimulus information is processed by the thalamus and then interpreted by visual association areas in the cerebrum. Extrinsic Eye Muscles Superior rectus • rotates eye up and medially Lateral rectus • rotates eye laterally Medial rectus • rotates eye medially Inferior rectus • rotates eye down and medially Superior oblique • rotates eye down and laterally Inferior oblique • rotates eye up and laterally THE END