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Unit 12: Evolutionary Processes Mr. Nagel Meade High School IB Syllabus Statements • 5.4.1 – Define evolution. • 5.4.8 – Explain two examples of evolution in response to environmental change; one must be antibiotic resistance in bacteria. • http://click4biology.info/c4b/5/eco5.4.htm Think… • What is meant by the word ‘evolution’? • What other words come to mind when you hear the word ‘evolution’? History of Understanding – Part I • The year is 1600… – You see what you believe to be fish coming from mud and flies from rotting meat – The accepted theory is spontaneous Spontaneous generation - living generation organisms can arise from non-living matter • How could you refute this notion in 1600? – Hint: The microscope hasn’t been invented yet! http://www.kent.k12.wa.us/staff/TimLynch/sci_class/chap01/spontaneous.html History of Understanding – Part II • Francesco Redi (1600’s) – Where do maggots come from? • Arise from spontaneous generation? • Rotten meat or fly’s eggs? History of Understanding – Part III • Lazzaro Spallanzani (1700’s) – Where do microorganisms come from? • Arise from ‘vital force’ in the air? – Nutrient baths » sealed and unsealed » boiled and not – Many people felt he ‘cooked the vital force out of the air’ History of Understanding – Part IV • Louis Pasteur (1800’s) – Where do microorganisms come from? • Final experiment to disprove ‘Vital Force’ – Unique glass shape traps dirt on which cells of organisms travel » Placed in various locations • Ended the era of ‘Spontaneous Generation’ • Began new era: Biogenesis! Biogenesis - theory that living organisms come only from other living organisms Charles Darwin • On the Origin of Species by Means of Natural Selection (1859) – Species originate through evolutionary change • HMS Beagle – Galapagos Islands (Finches) – 1838 [Ship:1831-1836] • http://www.bbc.co.uk/history/historic_figure s/darwin_charles.shtml Your Turn! - Create a Timeline • Include the following events, people, and ideas: – – – – – – – – – – – – Charles Darwin (HMS Beagle) – 1831-1836 Charles Darwin (Origin of Species) – 1859 Louis Pasteur’s Experiment – 1864 Lazzaro Spallanzani Experiment – 1767 Francesco Redi’s Experiment – 1668 Linnaeus (Classification system established) – 1735 Lamarck (Philosphie Zoologique) – 1809 Mendel’s papers published – 1866 Urey-Miller Experiment – 1952/1953 Scopes Monkey Trial – 1925 DNA puzzle solved – 1953 Human Genome project completed – 2003 Investigation: Adaptation • http://whyfiles.org/038badbugs/ – Read about how microbes have developed resistances to antibiotics • Germ Warfare • Putting up Resistance – How can adaptations influence an organism’s ability to survive? IB Syllabus Statements • 5.4.1 – Define evolution. • 5.4.5 – State that the members of a species show variation. • 5.4.6 – Explain how sexual reproduction promotes variation in a species. • 5.4.3 – State that populations tend to produce more offspring than the environment can support. • 5.4.4 – Explain that the consequence of the potential overproduction of offspring is a struggle for survival. • http://click4biology.info/c4b/5/eco5.4.htm Define • Evolution • Variation • Adaptation Vocabulary - Review • Evolution – A gradual process in which something changes into a different form • Variation – deviation of an individual from the group to which it belongs (the act or process of changing) • Adaptation – The adjustment or changes in an organism to become more suited to an environment Think…Pair…Share…Review • How can new traits come about (think sex)? – Result from new combinations of existing genes or from mutations of genes in reproductive cells within a population • How do adaptation and variation play into this concept? Natural Selection • The process by which organisms best suited to survival in their environment achieve greater reproductive success, thereby passing advantageous genetic characteristics on to future generations • How could changes in the environment bring about natural selection? Moths and Ind. Rev. • Gizmo? Survival of the Fittest • What consequences would result from having too many offspring? • What natural factors would ensure that the fittest organisms would survive? Warmup • What environmental pressures exist that naturally select members of a population to survive? IB Syllabus Statements • • • • • • • • • 5.4.2 – Outline the evidence for evolution provided by the fossil record, selective breeding of domesticated animals and homologous structures. D.1.1 – Describe four processes needed for the spontaneous origin of life on Earth. D.1.2 – Outline the experiments of Miller and Urey into the origin of organic compounds. D.1.3 – State that comets may have delivered organic compounds to Earth. D.1.4 – Discuss possible locations where conditions would have allowed the synthesis of organic compounds. D.1.5 – Outline two properties of RNA that would have allowed it to play a role in the origin of life. D.1.6 – State that living cells may have been preceded by protobionts, with an internal chemical environment different from their surroundings. D.1.7 – Outline the contribution of prokaryotes to the creation of an oxygen-rich atmosphere. D.1.8 – Discuss the endosymbiotic theory for the origin of eukaryotes. Video Clip • Darwin’s Theories in Action Brainstorm... • What evidences have you heard of that support evolution? • Evolution is a Theory. What is the difference between a theory and a law? Evidences of Evolution • Fossil Records – Transitional Fossils vs. Missing Links • Homologous Structures – It’s all Relative • Analogous Structures – Function, not Origin (Wings) • Vestigial Organs – Useful or Designer • Embryological Development – Fact or Fiction • Biochemical – AA sequences and Cytochrome C True Embryology Science 1997 Twenty years from now… • You are tasked with attempting to terraform Mars. – Research what factors were important in Earth’s early days to promote simple life. – What steps (and in what order) would you need to execute in order to achieve success? – What experiments/findings could substantiate your action plan? 4A’s plan for Mars Domination • Non-living simple organic molecules – Amino Acids (Methane, Ammonia, Water, H2) – Urey/Miller (presence electricity) • Polymerization – Heat/Energy (Volcanic activity, molten core) • Inheritance via self-replicating molecules – Ribozymes (RNA) • Package inside membranes – Prokaryotic (bacteria) eukaryotic (algae) Warmup • Name three subtypes of natural selection and provide an example of each! Warmup • What four processes were needed for the spontaneous origin of life on Earth? IB Syllabus Statements • D.2.1 – • D.2.2 – • Discuss ideas on the pace of evolution, including gradualism and punctuated equilibrium. D.2.10 – • Compare convergent and divergent evolution. D.2.9 – • Outline the process of adaptive radiation. D.2.8 – • Compare allopatric and sympatric speciation. D.2.7 – • Explain how polyploidy can contribute to speciation. D.2.6 – • Describe three examples of barriers between gene pools. D.2.5 – • Discuss the definition of the term species. D.2.4 – • State that evolution involves a change in allele frequency in a population’s gene pool over a number of generations. D.2.3 – • Define allele frequency and gene pool. Describe one example of transient polymorphism. D.2.11 – Describe sickle-cell anemia as an example of balanced polymorphism. Group Research • 1: Allele Frequency vs. Phenotypic Frequency • 2: Speciation and Adaptive Radiation • 3: Geographic vs. Temporal vs. Behavioral Isolation • 4: Allopatric vs. Sympatric Speciation • 5: Convergent vs. Divergent Evolution • 6: Gradualism vs. Punctuated Equilibrium • 7: Transient vs. Balanced Polymorphism Group Research • What is it? • Examples/Illustrations? • Why is this evolution? Warmup • What conditions are necessary for natural selection to occur? Definitions and Examples • • • • • • • • • • Gene Pool Allele Frequency Species Speciation Allopatric Speciation Sympatric Speciation Adaptive Radiation Convergent Evolution Divergent Evolution Equilibrium Allele Frequencies • How many people in class are male? Female? • How many X chromosomes are present? Y chromosomes? • Note: Chromosomes != alleles. • Large changes in allele frequency indicate evolutionary change! Species • Can interbreed and produce fertile offspring • Similar physiological and morphological characteristics (observed/measured) • Genetically distinct Speciation • Reasons for Speciation (mainly barriers) – Geographical • Allopatric [physical barrier] • Sympatric [shared area] – Temporal – Behavioral • Hybridization yields infertile offspring • Polyploidy – Incompatibility of chromosomal sets Adaptive Radiation • Adaptations yield variations on one species – Think Finches – Various adaptations allow a flavor of species to survive in their own environment – Over time, new species evolve (i.e. disruptive selection) Math • What do the terms converge and diverge mean? Physics? Equilibrium • Gradualism vs. Punctuated Polymorphism • Transient vs. Balanced Warm Up • Contrast panspermia with endosymbiosis. Warm Up • How does sexual reproduction promote variation within a species? IB Syllabus Statements • 5.5.1 – Outline the binomial system of nomenclature. • 5.5.2 – List seven levels in the hierarchy of taxa—kingdom, phylum, class, order, family, genus and species—using an example from two different kingdoms for each level. • 5.5.3 – Distinguish between the following phyla of plants, using simple external recognition features: bryophyta, filicinophyta, coniferophyta and angiospermophyta. • 5.5.4 – Distinguish between the following phyla of animals, using simple external recognition features: porifera, cnidaria, platyhelminthes, annelida, mollusca and arthropoda. • 5.5.5 – Apply and design a key for a group of up to eight organisms. • http://click4biology.info/c4b/5/eco5.5.htm Taxonomy • Linnaeus (1735) – (D) K P C O F G S Binomial Nomenclature • Bi = two – Genus (capitalized) – Species (not capitalized) • Should be italicized or underlined Plant Distinctions Phyla Bryophyta Filicinophyta Coniferophyta Angiospermophyta Features Animal Distinctions Phyla Porifera Cnidaria Platyhelminthes Annelida Mollusca Arthropoda Features Dichotomous Keys • Observed characteristics help divide species into smaller groups • Final step should have final identifying name Eight Species Develop a key that could be used to uniquely identify each organism, then give it to a classmate to verify that it works. Warm Up Develop a key that could be used to uniquely identify each organism, then give it to a classmate to verify that it works. IB Syllabus Statements • D.3.1 – • D.3.2 – • Discuss the correlation between the change in diet and increase in brain size during hominid evolution. D.3.9 – • Discuss the incompleteness of the fossil record and the resulting uncertainties about human evolution. D.3.8 – • State that, at various stages in hominid evolution, several species may have coexisted. D.3.7 – • Outline the trends illustrated by the fossils of Ardipithecus ramidus, Australopithecus including A. afarensis and A. africanus, and Homo including H. habilis, H. erectus, H. neanderthalensis and H. sapiens. D.3.6 – • Describe the major anatomical features that define humans as primates. D.3.5 – • Deduce the approximate age of materials based on a simple decay curve for a radioisotope. D.3.4 – • Define half-life. D.3.3 – • Outline the method for dating rocks and fossils using radioisotopes, with reference to 14C and 40K. Distinguish between genetic and cultural evolution. D.3.10 – Discuss the relative importance of genetic and cultural evolution in the recent evolution of humans. Radioactive Dating Techniques • Half-life: How long it takes for a substance to radioactively decay to half the quantity. • 14C (biomass) and 40K (rocks) are popular isotopes for determining age. • 14C = 5,730 yrs • 40K = 1.28 x 109 yrs • Proportion of stable daughters to unstable parent isotopes How old is it? • Given the curve above, how old is the sample (tested for 14C) if the sample has roughly 6% 14C remaining in it? • 14C = 5,730 yrs • Amount = 1 / (2t½) Research Time! • Outline the trends illustrated by the fossils of: • Ardipithecus ramidus • Australopithecus – A. afarensis – A. africanus • Homo – – – – H. habilis H. erectus H. neanderthalensis H. sapiens. Anatomical Differences Anatomical feature Ape-like form Human-like form Position of spinal intersection with skull Towards the back of the skull base Towards the center of the skull base Cranial Capacity Small Large Canine Teeth Long/Sharp Short/Dull Molars Long/Narrow Short/Wide Brow Ridge Protruding Flat Facial Region Protruding Flat Jaw Tall/Thick Small and Thin Adapted from HL Biology page 436 Hominid Evolution I • Uncertainties: 5 million years of evolution with only a few ‘pieces’ in the puzzle. • Doubtful that any species interacted with each other, even though some coexisted on Earth at the same time. (Carbon dating) • Increased proteins/meats in diet led to increased brain size/energy requirements. • DNA implies a common ancestor with apes, but no scientist has ever proposed being DESCENDED from monkeys. (radiation) Genetic vs. Cultural Evolution • Genetic Evolution implies changes in INHERITED CHARACTERISTICS – Morphology – Chromosomal Numbers – Biochemistry (blood proteins, enzymes) • Cultural Evolution implies changes in ACQUIRED KNOWLEDGE – – – – Language Customs and Rituals Art Technology Hominid Evolution II • As brain size increased, quality of tool making also increased. • Genetic evolution precedes cultural evolution. • In the last 30k years, H. Sapiens have been largely cultural. – Is this atypical? Warm Up • Describe the importance of changes in allele frequency for the evolution of one species into another. (4 marks) natural selective pressures result in survival of advantageous alleles; frequency of these alleles will increase through reproduction; these alleles spread through population; basis for microevolution; over time many advantageous genes accumulate in a species; when many changes occur some members of a species cannot successfully mate with others / reproductive isolation; results in evolution of a new species; IB Syllabus Statements • D.4.1 – • D.4.2 – • D.4.3 • D.5.1 – – • • D.5.7 • D.5.8 – – Discuss how biochemical variations can be used as an evolutionary clock. Define clade and cladistics. Distinguish, with examples, between analogous and homologous characteristics. Outline the methods used to construct cladograms and the conclusions that can be drawn from them. Construct a simple cladogram. D.5.9 – • Explain how variations in specific molecules can indicate phylogeny. D.5.6 – • Explain the biochemical evidence provided by the universality of DNA and protein structures for the common ancestry of living organisms. D.5.5 – • Outline the value of classifying organisms. D.5.4 – • State the assumptions made when the Hardy–Weinberg equation is used. D.5.3 – • Calculate allele, genotype and phenotype frequencies for two alleles of a gene, using the Hardy–Weinberg equation. D.5.2 – • Explain how the Hardy–Weinberg equation is derived. Analyse cladograms in terms of phylogenetic relationships. D.5.10 – Discuss the relationship between cladograms and the classification of living organisms. Hardy-Weinberg Warm Up Warm Up Answers