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Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Concepts and Methods “the Big picture” organization cell organelle organism organ system family population community, ecosystem, biosphere producers, consumers, decomposers life and death unity in life diversity of life interdependency homeostasis growth energy metabolism DNA, RNA, reproduction, heredity Mutation/evolution adaptation Diversity Classification/taxonomy Animals Plants Fungi Monerans: two kinds protists kingdom, phylum, class, order, family, genus species Theory Scientific Method Hypothesis Conclusion control group data bias, error, accuracy and precision Practical: Every moment of your life you are constantly involved with life – your own, those of others, those of symbionts and of sources of food and materials. Death of many organisms is required to support your life. Homework: Do the self quiz on your own and be prepared to answer the Review Questions: page 17. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Chemical Foundations Unity of life concept, revisited Atoms subatomic particles: proton electron neutron atomic number atomic mass isotopes radioactivity (decay) electron orbitals/shells molecules mixtures compounds bonding: covalent ionic - disassociation hydrogen hydrophilic/ hydrophobic substances reactivity solvent, solute, solution, solubility pH acid/base buffer salt mineral Practical Application: A basic understanding of chemistry is fundamental in many of your daily activities. Organic chemistry is based on a small set of elements and is the basis of life, nutrients and medicines. Your understanding of hormones and toxic substances, especially carcinogens and mutagens is enhanced by a basic appreciation of life chemistry Homework: Get ahead in your study of Biology by reading the next assigned chapter. Continue to do the self quizzes on your own. Also study the review questions and be prepared to answer them. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Carbon Compounds in Cells Organic compounds Bonding behavior Tetrahedron Synthesis (anabolism and catabolism) Functional groups: Hydroxyl Aldehyde Ketone Carboxyl Amino phosphate Chains and rings Classes of reactions Functional group transfer Electron transfer Rearrangement Condensation/dehydration Cleavage/hydrolysis Monomers and Polymers Saturation, unsaturation and double bonds carbohydrate mono, di, oligo and polysaccharides starch cellulose glycogen lipid, hydrocarbons, nonpolar fatty acids triglycerides phospholipids sterols protein, polypeptides amino acids disulfide bridge levels of structure: primary, secondary, tertiary, quaternary glycoprotein denaturation nucleic acid DNA, RNA Nucleotides Riboses Phosphate 4 Bases (G, C, A, T) Double/single helix ATP Homework: Get ahead in your study of Biology by reading the next assigned chapter. Continue to do the self quizzes on your own. Also study the review questions and be prepared to answer them. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Cell Structure and Function Cell theory Every organism is composed of cells The cell is the smallest life unit All life comes from living cells Structural organization/compartmentalization Plasma (cell) membrane Cytoplasm Cytoskeleton Nucleoid region/nucleus: prokaryotes/eukaryotes Cell membrane Lipid bilayer, phospholipids Micelles Surface-to-volume ratio Microscopy Light, TEM, SEM Micrograph Eukaryotic Organelles Nucleus, envelope (pores), nucleolus, chromosomes chloroplast mitochondria cytomembrane system: endoplasmic reticulum (ER), smooth and rough (ribosomes) golgi body vesicles, lysosome central vacuole centrioles, basal bodies Cytoskeleton components Microtubules Microfilaments Intermediate filaments Motility Pseudopods Flagella and cilia (9+2 array), MTOC Streaming Specialization Cell wall Matrixes Plasmodesmata Prokaryotic structures Membrane Nucleoid Ribosomes Cell wall Capsule Pili Bacterial flagella Practical Application: To know the parts of cells and their functions is to begin to see how tissues provide functions in organs. Also the way a disease organism affects the body is largely understood at this level. Homework: Get ahead in your study of Biology by reading the next assigned chapter. Continue to do the self quizzes on your own. Also study the review questions and be prepared to answer them. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Cell Membranes Phospholipid Cholesterol fluid mosaic lipid bilayer diffusion solute, solvent, solution gradient concept concentration electric pressure membrane proteins (including glycoproteins) transport open and gated channel proteins (passive transport, facilitated diffusion) active transport (pumps, ATP) reception recognition adhesion endocytosis receptor mediated, bulk phase, phagocytosis exocytosis, pinocytosis (contractile vacuoles) vesicles and membrane cycling, golgi apparatus, lysosomes osmosis passive diffusion of water concentration of solute iso, hypo, and hypertonic plasmolysis Practical Application: Reverse osmosis is a practical way to produce pure water from salty or contaminated water. Understandings of diffusion, osmosis and active transport explain a variety of biological phenomena including nervous impulse conduction, wilting and sap flow in plants, venom concentration in snakes and spiders and electrical discharge in eels. Homework: Start getting ready for the first exam. Study terminology and look for likely essay questions in your readings. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Metabolism Metabolism Anabolism/Catabolism Energy Potential, Kinetic, Thermal (heat) Molecular potential (chemical) Calories Chemical Reactions Laws of thermodynamics 1. Total energy remains constant 2. Entropy increases (3. life exists) Law of the conservation of mass reactants -> products concentrations directional equilibrium concept (bi-directional) exergonic/endergonic reactions Energy transfer ATP, ADP, phosphorylation Coupled reactions Metabolic pathways Degradative/Biosynthesis Substrates/intermediates/end products Energy carriers Enzymes and Cofactors Transport proteins Enzymes Catalytic Reused Bidirectional Selective Mechanism: Activation energy concept Active sites Induced fit concept Transition state Environmental factors: Temperature pH feedback inhibition coenzymes Electron transfer and transport systems Oxidation-reduction reaction Electron donor, Electron acceptor, Pathways of enzymes/proteins Practical: This is life whether we want to acknowledge it or not. These are the mechanisms God put in us to make us and other life forms function in the material world. The proteins of life are the cogs, machines and building blocks. ATP is the fuel. Much of the future of Biology as a field of study will focus on proteins. Homework: Start getting ready for the first exam. Study terminology and look for likely essay questions in your readings. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Energy Acquisition Photosynthesis/Aerobic respiration Photoautotroph/heterotroph Light Electromagnetic radiation Spectrum Visible light Wavelength Prism Pigments Absorption spectrum Chlorophyll a and b Fluorescence Accessory pigments: caratenoids, antho and phycobilins Photosystems: I and II Light harvest Reaction centers: electron transport systems, ATP formation (noncyclic) Light dependent reactions Chemiosmotic theory of ATP formation Excited electrons Photolysis of water (H+ and Oxygen gas) ATP or NADPH formation Thylakoid membranes of chloroplasts Light-independent reactions (dark) Carbon fixation C-3, C-4 and CAM Calvin-Benson cycle RuBP, PGA, PGAL, glucose ATP and NADPH requirement Starch Cellulose Practical application: Think about the environment when you contemplate the concepts explained in this chapter. Think about the effects that photosynthesis has on the chloroplast, he leaf cells, the leaf vessels and on the air surrounding the leaves. There are a number of connecting surfaces involved that give an interface between the inner cellular photochemistry and the outside environment. We, as air-breathers, also interface with this same air. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Energy-Releasing Pathways ATP formation from compounds made by producer organisms Aerobic Respiration Glycolysis (in cytoplasm) Glucose Phosphorylation Pyruvate ATP input/output, (net) Can be part of anaerobic respiration: see below Krebs Cycle (in mitochondria) CO2 produced ATP formed Coenzyme A, acetyl-CoA Citrate, isocitrate, alpha-ketoglutarate, succinyl-CoA, succinate, fumarate, malate, oxaloacetate NADH, FADH2, ATP Electron Transport System (between compartments of the mitochondria) H+ formed as NADH and FADH2 deliver electrons to the system O2 accepts electrons, makes H2O with acid ATP formed as H+ reenters the inner compartment Anaerobic respiration= fermentation Glycolysis Same pathway Will stop if pyruvate builds up Fermentation Pyruvate converted to lactate to accept electrons Ethanol formation by way of acetaldehyde CO2 released Alternative pathways Food digestion Intracellular conversions Fats Glycerol becomes PGAL Fatty acids become PGAL Carbohydrates Glycogen Proteins Aminoacids carbon backbones for pyruvate and acetyl-CoA amino (ammonia) dumped as Urea Practical applications: The understanding of these metabolic processes is the basis of many specialty areas in animal and human science, including nutrition, digestion, biosynthesis, physiology, muscular energetics, thermoregulation and the production and excretion of nitrogenous wastes (kidney function). In industry these processes are so well understood that many important microorganisms are practical to grow to process for extracting including a wide variety of organic compounds and food additives. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Mitosis Cellular reproduction Growth Division/fission Multiplication Mother cells -> daughter cells Inheritance, duplication, distribution, mechanisms Nuclear phenomena Non-nuclear considerations Chromosomes Chromatid: Sister chromatids: Centromere Ploidy Diploid: 2n Haploid: 1n Other ploidy: tetraploid Mitosis: Somatic cells 2n -> 2 x 2n -> 2n + 2n Meiosis: (reduction of diploid to haploid) germ cells 2n -> 2 x 2n -> 2 x 1n + 2 x 1n -> 1n + 1n + 1n + 1n Cell cycle G1 = first growth period S = DNA synthesis period G2 = second growth period M = mitosis Prophase Metaphase Anaphase Telophase Cytoplasmic division/Cytokinesis/cleavage or cell plate formation Components Condensed chromosomes Chromatin Histones Nucleosomes Kinetochore Spindles 10k microtubules motor proteins cell plate (plants) vesicle convergence Practical Applications: Knowledge of mitosis and the cell cycle is the basis of organismal growth, vegetative (asexual) reproduction, tissue healing, regeneration of lost body parts and cancer. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Meiosis Sexual reproduction Gametes sex cells sperms eggs (ova) spores Gametogenesis Germ cells Spermatogenesis Oogenesis Fertilization, zygote formation, conception Chromosomes, sister, chromatids, homologues Ploidy: diploid haploid the “meaning” of the diploid numbers of organisms genes and alleles karyotype, genotype and phenotype Stages of meiosis Meiosis I Prophase I Metaphase I Anaphase I Telophase I Cytokinesis Meiosis II Prophase II Metaphase II Anaphase II Telophase II Cytokinesis Critical alignments Reduction Crossing over, recombination and chiasmata Alternation of generations Practical Applications: Knowledge of meiosis is the basis of understanding sexual reproduction. Mendel’s laws are best understood in view of what occurs in meiosis. Odd forms of meiosis lead to diseases that are not uncommon in our species. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Observable Patterns of Inheritance Genetics Genes, loci, homologous chromosomes, alleles, pairs Inheritance “law” of blending Mendelian genetics True (pure) breeding/hybrids Homozygotes/heterozygotes Dominant/recessive traits Genotype/phenotype P, F1, F2 Crosses: mono and dihybrid, test crosses Punnett squares and probability Laws Segregation Independent assortment Modern understandings Chromosomal linkages Dominance varitions Incomplete Codominance Pleiotropy Interactions: epistasis Polygenetics Continuous variation Environmental effects Extranuclear inheritance Practical Applications: Knowledge of basic genetics explains most of what happens in the expression of the genotype into a phenotype. Don’t forget “Nature vs. Nurture”. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Chromosomes and Human Genes Chromosomal basis of inheritance Gene, chromosome, homologues, recombination, meiosis, gametogenesis Autosomes and sex chromosomes Karyotypes analysis Sex determination in humans X and Y chromosomes Development Linkage groups X and Y linked genes, reciprocal test crosses Recombination Tightness of linkage Length of chromosome Mapping Map units = frequency of recombination Physical units Human genetic analysis Pedigrees Symbols Patterns Genetic disorders Autosomal Recessive Dominant x-linked dominant recessive nondisjunction euploidy, aneuploidy and polyploidy autosomal: Down sex chromsome syndromes: Turner, Klinefelter, XXY, XYY chromosome structure abnormalities deletion, duplication, inversion, translocation age of onset of effects of disorder treatments, counseling, screening, prenatal diagnosis Preimplantation diagnosis Practical Applications: To be forewarned of an impending birth of a genetically different child is to begin to prepare the whole family for the special accommodations that will be needed. Abortion is not an option. Homework: do the odd problems and check your work. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes DNA Structure and Function “This molecular constancy and variation among species is the foundation for the unity and diversity of life.” History 1868 – Miescher: puss and semen studies led to isolation of nucleic “acids” 1928 – Griffith: mouse studies of “dead” disease traits inheritance 1949 – Chargraff: [Adenine] = [Thymine] and [Guanine] = [Cytosine] 1951 – Pauling: deduced 3D structure of protein 1951 – Franklin: x-ray diffraction images of DNA (compared to known chemicals) implied helical structure 1953 – Watson and Crick: proposed molecular structure of DNA 1950’s – Delbruck, et al, studies of bacteriophage DNA cores DNA Nucleotides: one base to one phosphate to one deoxyribose Helical structure Double/ two-stranded 3.4 nm per full twist opposing directions – bidirectional base pairing – hydrogen bonds base sequence a “sense” strand the order of G, C, A, T self replication semiconservative: one of the old strand resides in the new pair of strands enzymes: DNA polymerase, ligase, unwinding enzymes replication forks: origins at binding sites, bidirectional, triphosphorylated nucleotides create new strand continuously discontinuously: Okazaki fragments Chromosomal proteins: scaffolding between genes? DNA repair Excision repair UV light Cancer and mole plotting Practical application: The understanding of DNA replication is at the heart of the cellular reproduction of life. The success of our built-in DNA repair mechanisms ensures a near perfect continuity of life. It also explains how some damaged cells survive to give us cancers and other mutated tissues. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes From DNA to Proteins Garrod’s hypothesis: inherited trait abnormally high levels of urinary (metabolic) substances related to a key enzyme in a metabolic pathway One gene – one polypeptide: Transcription and translation required RNA Three kinds mRNA = Messenger = gene transcript – made as needed tRNA = Transfer = has RNA “anticodon” and carries AA’s – made abundantly for all translations rRNA = Ribosomal = special transcript to make new ribosomes – takes one to make one Transcription RNA assembly DNA unwinding Role of uracil: A – U not A - T Promoter RNA polymerase DNA rewinding Processing (modification of transcript) Introns and exons Excision Poly-A tail Translation Migration of mRNA through nuclear pores into ctoplasm Codons – the genetic code Base sequence triplets 64 translate to 20 AA’s Ribosomes Two kinds Assemble in ER Sites of mRNA translation and polypeptide assembly tRNA Possess anticodon Enzymatically attached to appropriate AA Transfer the AA to ribosome and enter in according top the mRNA sequence Stages Initiation mRNA loaded into ribosomal unit – AUG = START initiation complex formed appropriate tRNA locked in Elongation Polypeptide chain forms and grows Sequence of AA’s determined by codons on mRNA Forming polypeptide begins to take on its 3D form mRNA may enter another ribosome to start the translation process again before completing the first translation – chains of ribosomal events Termination STOP codon halts elongation Release factors Final protein “shipped out” Gene mutations Simple mutation Base-pair substitution Insertion Deletion Complex Transposable elements Causes: spontaneous, mutagenic chemicals and ionizing radiation Reverse translation?? Practical application: This is much of what complex life at the whole organism level amounts to: genes in active cells decoding into mRNA’s to become proteins and expressed traits. Development is related to the polarity, the environment and the intercellular communication networks that form in the embryo to control/regulate these processes.. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Controls Over Genes Genetic expression – cells exert control over selves and each other the phenotype cell type and function cell environment – chemicals, signals and outside factors development adaptation programmed cell death control systems concept regulatory proteins – interactions operon concept: promoter, repressor binding site, linked genes prokaryotic controls: simple, on-off, rapid response to environmental changes negative control repressor compound blocks promoter (target molecule) inhibition of transcription presence of a signal chemical removes repressor positive control promotion of transcription activator protein eukaryotic controls: much more intricate in complex organisms cell differentiation embryonic origins specialization cell activity: selective expression transcription controls gene amplification DNA rearrangements Chemical modifications post-transcription controls transcript processing: alternative splicing translation controls: degradation and inactivation protein processing: activation, inhibition, stability short and long term aspects of expression Examples Transcription: lampbrush chromosomes observed in amphibians X chromosome inactivation Barr bodies and skin mosaic Calico cats Signaling mechanisms Hormones Stimulation or inhibition of target cells Enhancers – binding sites Hormonal control of gene expression is like a full symphony orchestra Sunlight signals Phytochrome in plants Cancer revisited Oncogenes Loss of gene control Growth factors Tumors: benign vs. malignant Metastasis Traits of cancer cells Changes in membrane and cytoplasm Abnormal rate of cell division Weakened adhesion lethality Practical applications: This is what genes are all about. Understanding these gene expression control mechanisms is the first step in understanding the mystery of life. These are also the “tip of the iceberg” in the whole story. We do not yet know all of the intricacies of the interactions involved. Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Recombinant DNA and Genetic Engineering Germplasm Technology, engineering and therapy Sports and hybridization DNA Restriction enzymes RFLP’s (restriction fragment length polymorphisms) Modification enzymes DNA amplification Cloning vector - plasmids Polymerase chain reaction (PCR) - billions quickly cDNA and reverse transcription based on mRNA DNA fingerprinting Tandem repeats Gel electrophoresesis DNA sequencing Gel electrophoresesis DNA libraries Probes Screening Application Recombination –a new definition Plant tissue culture (an aside) DNA ligase Recombinant plasmids Genetic alteration (engineering) Beneficial genes Methods of gene transfer Plasmids Viruses Electroporation Bacteria Plants Animals Human genome project 35,000 have been studied so far 0.1% of human genome varies = 3,200,000 bp 400 are related to genetic disordered Gene therapies in humans Human enhancement Safety issues Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Emergence of Evolutionary Thought “In the beginning…” “Any population can evolve when individuals differ in one or more inheritable traits that are responsible for differences in the ability to survive and reproduce.” Biological Science History lesson – a quest for the “meaning of life” and the “secret of life” Ancient Greece Hippocrates -Natural Cause theory Aristotle – Continuum of organization, taxonomy of kinds Early Christians Genesis, the inspired word of God Creation of species The Garden The Fall European exploration of the world Biogeography data first compiled Overwhelming quantities of information Many kinds Wide distribution Similar traits patterns Center of creation theories (where was the garden?) Comparative morphology, anatomy and physiology Patterns of similarity Basic body plan theory Homologous organs Snakes have leg bones! Geology - fossils Sequential prehistoric data Multiple sites of origin theories Newer species descending from older theories (evolution) Global awareness – 1900’s Rock-solid Christian Faith: God is the designer and distributor of Earth species Emergence of modern Christian explanations of God’s creation Atheistic explanations and challenges – Is God dead? Theories emerging Cuvier – mass extinction and catastrophism theory Lamarck – environmental factors and the “acquired characteristics” theory Lyell - Theory of uniformity – Wegener – continental drift Darwin and Wallace – natural selection theory Traits Survival adaptations What does “Hindsight is 20-20” mean in this context? Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Microevolution Population concept of individuals with traits morphological physiological behavioral Gene pool of alleles (remember how genes are expressed and controlled) Polymorphism Continuous variation (small, incremental differences in genotypic expression) Inheritance: 10600 combinations possible Two parents (each 223 chromosomal combinations) Independent assortment Crossing over Mutation (gene deactivation) Chromosomal abnormalities Allelic frequencies Hardy-Weinberg rule Genetic equilibrium (an idealized case: a kind of theory) No mutation Large population Gene pool isolation (no immigration) Gene not repro or survival critical Random mating Changes caused by circumstances Microevolution: small scale changes in the gene pool Gene flow Natural selection (viv a vis artificial selection) Genetic drift Mutation revisited Rate: 10-5 per gene locus per gamete per generation one gamete per 100,000 is mutated Lethal Neutral Beneficial ???? Natural selection Reproductive capacity Limited resources/competition Gene pool concept Variation in phenotypes Fitness: according to current environment Natural selection results Kinds Directional: resistances Stabilizing: fixing traits Disruptive: splitting the gene pool Balancing: sexual dimorphism Drift Bottle necks Founder effect Speciation Practical application: Was there speciation? Is it occurring now? Where? Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Speciation “Species are groups of interbreeding natural populations that are reproductively isolated from other such groups.” Species (kind) Morphological definition: based on appearance life stages questions effects of environment can morphologically identical organisms be different species? Biological definition: applies to sexually reproducing organisms Based on reproduction Same as long as form, physiology, behavior permit interbreeding Fertile offspring Origins? That is the question? Genetic change and isolation (speciation) Not purposeful, a byproduct of genetic change Gene pools’ gene flows and divergence Gradual and variable in pattern Mechanisms of reproductive isolation and genetic compatibility Prezygotic Temporal Behavioral Mechanical Ecological Gametic mortality and biochemical changes Postzygotic Early death of embryo Implantation failure (in mammals) Offspring inviability Offspring sterility Role of geography (allopatric speciation) Barriers Time factors Drift Founder effect and bottlenecks Other kinds of speciation Sympatric Polyploidy Parapatric (sketchy evidence) Two main patterns Cladogenesis: branching (two from one) Anagenesis: new from old Diagrams: Trees Gradual Punctuated Adaptive radiation concept Extinctions Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes The Puzzle Don’t forget what it says in Genesis Microevolution vs. Macroevolution Macro patterns: long term studies of lineages Genetic persistence: basic unifying traits of life Genetic divergence: speciation Genetic disconnect: extinction Fossils: 250,000 species? Fossilization Imprints Stratification: Law of Superposition Concept of a “fossil record” Gaps and completeness Readability? Time Biblical Geologic Time scale (mya) Eras, periods, epochs Plate tectonics Continental Drift: 1908 – F. Taylor Pangea – Alfred Wegener Seafloor spreading Magnetic field alignment Comparative Biology Embryology Theory:“The early embryos of vertebrates strongly resemble one another BECAUSE they have inherited the same ancient plan for development” Morphology Homologous structures in widely divergent species Analogous (similar) structures in (seemingly) unrelated species – convergence Biochemistry Genetics Close genetic similarities between species with pronounced morphological differences Molecular clock concept Proteins AA sequences Taxonomy and Systematic Biology Binomial naming system: Genus and species Kpcofgs Sub and super groupings added Developed from lower to higher Phylogeny now is a directing force Cladistic taxonomy Traits quantified Cladograms How many kingdoms Whittaker – 1969 – five Archaebacteria – sixth kingdom Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Theories of the Origin of Life Why connect this to our current studies? “Although the story is not yet complete…” “There are major gaps in the story of life’s origins.” Questions (always remember Genesis: In the beginning…) What was Earth like when life first appeared? Could it have originated spontaneously? Theories? Must it have come from off world? Theories? Can we devise experiments and computer models to test the plausibility and support or reject these? Tools Curiosity, logic and “lateral” thinking (imagnation) Lenses: Telescope, Microscope Sciences: Biology, Chemistry, Physics Faster, more capable computers Evidences Time Earth Materials: atmospheric gasses, crust, mantle, oceans, salts and other dissolved minerals Astrophysics Other planets atmospheres : was the first Earth atmosphere oxygen free? Planetary origin theories Miller’s experiments Fox’s studies Origin of life processes Metabolic agents, catalysts? Self replication: RNA first? DNA? Connection to protein synthesis? Membranes Containers? Osmosis? Electrical potentials Active transport? Micelles Proto-cell concept Stromatolites – mat fossils A Tree of lineages Anaerobic origins of three life types (Archaen Eon) Eubacteria Archaebacteria – eventually three kinds emerge Megacells Photosynthesis originated in eubacteria Atmospheric oxygen increases (Proterozoic Eon starts) Endosymbiont concept Engulfed bacteria as organelles Megacells host certain kinds of eubacteria – first eukaryotes – animals emerge Some of same kinds of cells host photosynthetic eubacteria – plants emerge Atmosphere becomes more oxygenated (20%) Land organisms emerge from the sea Major events theoretically displayed in the fossil record Continental migration Mass extinctions Subsequent radiations Practical application: Does the high diversity and distinct kinds distributed over the continents reflect any natural phenomena at play or is it all according to a detailed original plan of God’s design? Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Ch 49: The Biosphere Ecology – levels of study: Earth scale: Biosphere level – interactions with lithosphere, hydrosphere, atmosphere Climate: Temperature zones: Latitude Altitude Precipitation zones: air and water circulation patterns Ocean currents Prevailing winds Topography: rain shadow example Biomes 6 terrestrial biogeographic realms Predictable arrangement of biomes Soils Specific biomes Desert Shrublands, woodlands and grasslands Tropical forests Coniferous (boreal) forests Tundra: arctic and alpine Hydrosphere level Freshwater provinces Lakes Zonation: littoral, limnetic, profundal Overturn and the thermocline Tropic conditions Streams Riffles, pools and runs Ocean provinces Plankton and marine “snow” Hydrothermal vents LME’s – large marine ecosystems (ecoregions, ecozones) Benthic vs. pelagic Coastal: interface of land and sea Coral reefs Mangrove Estuaries Intertidal zone: rocky vs. sandy Upwellings El Nino Faulkner University Principles of Biology (BIO 1401) Science Department Chapter Notes Ecosystems Structure and function are similar in all ecosystems Trophic levels: “Who’s who” = who eats whom Plants rule (primary producers) Herbivores - primary consumers Carnivores – secondary consumers Detritivores and decomposers Other vores Omnivores Insectivores Granivores Frutivores Energy flows (one way) through the ecosystems: Light Carbon compounds Heat Light again Food chains and webs Ecological pyramids Gross and net primary productivity – kcal/sq m/yr Harnessing solar energy Transfers and “losses” Biogeochemical (Nutrient) cycles Cycling not flowing Hydrologic Carbon Nitrogen Phosphorus Modeling Greenhouse effect and global warming Faulkner University Principles of Biology (BIO 1401) Ecosystems (continued) Ecology at the Community level Niche concept: role in the “play of life” specific means of acquiring food Habitat, territories and ranges Food chains Food webs Symbiotic relationships Predation One way benefits predator/prey Parasitism One way benefits Host/parasite External and internal Parasites of parasites Commensalisms One way benefits Internal and external Mutualisms Two-way benefits External and internal Families Human parallels Carve out “your own niche” Interpersonal relationships Career choices – societal roles Science Department Chapter Notes