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17-4: Patterns of Evolution What an organism eats and does (physical and biological conditions), and where it lives in its environment is a NICHE 2 species that occupy the same niche create COMPETITION Competition can lead to EXTINCTION EXTINCT MACROEVOLUTION Definition: Large scale evolutionary changes that take place over long periods of time. Six patterns of macroevolution 1. Mass extinction (read and know!) 2. Adaptive radiation 3. Convergent evolution (analogous structures) 4. Coevolution (read and know!) 5. Punctuated equilibrium 6. Changes in developmental genes (read and know!) A Example: Galapagos Islands (Darwin’s Finches) B S. America Process of one species giving rise to many species that live in different ways (niches) AKA: divergent evolution Organisms evolve a variety of characteristics that enable them to survive in different niches Hawaiian Honeycreeper Convergent Evolution • Different organisms (unrelated) look similar because they live in similar environments • Different “raw material” for natural selection to work on, but… – Similar environmental demands • EX: moving through air, water, eating similar foods Convergent Evolution • Produces analogous structures like the dolphin’s fluke and a fish’s tail fin – Look and function similarly but do not share a common evolutionary history Darwin’s idea of evolution • Darwin thought evolution only took place over a LONG time – Hutton and Lyell’s discussion of slow geologic change • gradualism = fossil record continuous minor changes (Evolution is slow and steady!) Punctuated Equilibrium • Equilibrium—hardly any change • Definition: A pattern of long stable periods interrupted by brief periods of rapid change Examples • When the equilibrium is upset, change can occur in a short period of time • EX: A small group of organisms migrate to a new environment – Organisms evolve quickly to fill available niches (Galapagos Finches) • EX: A small population is cut off from its original population Example of Punctuated Equilibrium • Life is going on smoothly for a population of mice. • Then whoosh! • There is a flood which separates the population into two groups, one on one side of a river and one on the other side. (Geographic isolationreproductive isolation!) • What could happen as a result? Developmental Genes Developmental Genes “master control genes” = Hox gene controls embryonic development in animals controls what turns into the front or back part of animal and what becomes the arms, legs, etc.) Hox genes can reveal how evolution occurred (through molecular studies) 1. Provide the same body plan in all animals 2. Major evolutionary changes may be based on hox genes (example: # of wings, legs, body segments) Developmental Genes 3. Small changes in the timing of this genetic control during development can affect the body type of the organism (long legs vs. short legs) THUS contributing to the variation involved in natural selection. example: small change in when gene turned “on or off” would determine long vs. short legs; 3 vs. 4 wings THIS IS A BIG FIELD OF RESEARCH— UCLA has great evolutionary biology department.