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					Origin and Diversification of Eukaryotes: Protists Eukaryotic Cells  Nucleus and membrane bound organelles  Well developed cytoskeleton – structural support that lends to asymmetric forms  May have originated more than 2.7 bya, but 1.8 bya widely accepted 3 stages of diversity documented by fossil records  Initial diversification – 1.8 – 1.3 bya  Origin of multicellularity 1.3 bya – 635 mya  Emergence of large eukaryotes 635-535 mya endosymbiosis  Symbiotic relationship in which one organism lives inside the bosy or cell of another organism.  DNA sequence data suggest that eukaryotes are “combination” organism from Archaea and Bacteria  Endosymbiont theory – mitochondria and plastids were formerly small prokaryotes that began living within larger cells. (See page 485) Multicellularity  Colonies – collection of cells that are connected but show little or no cellular differentiation.  Multicellular organisms with differentiated cells – Algae, plants, fungi and animals Characteristics  Most diverse kingdom        eukaryotic., 1.5 bya Primarily unicellular/multicellular, heterotrophic/autotrophic Usually asexual, some sexual Found in water, damp soil and sand, leaf litter Some are parasitic 14 phyla Characteristics  Characteristics first seen in Protista kingdom    Sexual reproduction (varies, mitosis and meiosis – usually in harsh enviro.) Multicellularity (coordination among specialized cells) Complex flagella and cilia (as opposed to simple seen in bacteria) Protist diversity  Heterotrophic protists = protozoans, slime molds, water molds, parasites  Photosynthetic protists = Algae  Some are classified further by how they move:    Amoebas – pseudopodia Ciliates – cilia, ex. paramecium Flagellates – flagella, ex. Euglena Algae – Green, Red and Brown  Strict autotrophs, some multicellular  Distinguished by type of pigment they contain  Red  Multicellular, warm ocean waters  Red pigment can absorb light in deep water  Brown  Multicellular, marine  Kelp, grows along coasts, food and shelter  Green  Most freshwater, uni and multicellular  Contains chlorophyll a and b Green Algae  Phylum Chlorophyta      Most unicellular, can be filamentous or colonial Chlamydomonas – unicellular green algae Spirogyra – filamentous green algae Ulva – multicellular green algae (sea lettuce) Volvox – colonial (loose association of independent cells) green algae Red Algae/Brown Algae  Phylum Rhodophyta  Economically important –    Agar Wrappings around sushi Emulsifying agent for production of chocolate  Phylum Phaeophyta  Contain chlorophylls a and c and carotinoid  Kelp, rockweed – grow along shoreline  Harvested for human food and fertilizers  Algin – pectin-like material added to icecream Diatoms  Phylum Chrysophyta – includes golden-brown algae and yellow-green algae  Photosynthetic, significant part of phytoplankton, Important to food chain  Double shell made out of silica, like hat box  Move by excreting chemicals through Diatomaceous earth – abrasives  holes in shell   Found in oceans and lakes Dinoflagellates  2 flagella, cellulose plates  Unicellular  Most are marine, make up part of the plankton, important food source, producers  Some produce powerful toxins – “red tide” Euglena  Freshwater, unicellular with 2 flagella  Some photosynthetic with chloroplasts and has ability to ingest food as well  Pellicle – protein fiber inside cell membrane, allows for flexibility and shape change  Eyespot – light sensitive organ, toward light  Contractile vacuole – expels excess water Zooflagellates  Mostly unicellular, hetertrophic protozoans  Symbiotic and parasitic forms  Trypanosomes – transmitted by tsetse fly  Causes African Sleeping Sickness  Giardia lamblia – cysts from contaminated water, intestinal wall, severe diarrhea  Trichomonas vaginalis – STO, infects vagina and urethra of women, prostate of males Amoeba  Pseudopodia – false foot, cytoplasmic extensions  Food vacuole – food enters and moves throughout cell  Contractile vacuole – regulates water  Entamoeba histolytica – amoebic dysentery Forams  marine protist that lives in sand or attaches to other organisms  Tests – porous shells  Thin projections of cytoplasm extend through pores and aid in swimming Paramecium  Freshwater streams and ponds  Unicellular, heterotrophic  Cilia to move  Two nucluei   Macro – cell activities, includes DNA Micro –cell reproduction, contains chromosomes Sporozoans – parasitic protists  Form spores during their reproductive cycle  Nonmotile, spore forming, unicellular parasites  Malaria – anopheles mosquito, pg 392  Toxoplasma gondii – toxoplasmosis, during pregnancy, can affect fetus Slime molds - decomposers  Cellular slime mold, small    Behave like amoebas Ingest bacteria During stress, form multicellular colonies  Plasmodial slime molds   Mass of cytoplasm that looks like ooze, colorful Eat bacteria and other organic material, phagocytes Water Molds  Filamentous decomposers  Parasitic to plants and animals in ecosystems  Cell wall contains cellulose, not chitin  Usually have furry growths  Responsible for potato famine in Ireland, 1840’s Beneficial protists  Commensal protists that live in digestive tracts of humans and livestock (hay, cellulose)  Plankton – food, shelter, producers  Largest group of photosynthesizers  Abrasives  Detritivores – recycle chemicals in environment  Symbiotic relationships - coral
 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
									 
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                             
                                            