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Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ Section I: True or False statements. (½ mark each - no penalty for wrong answers). Q Statement True 1 A basic tenet of the theory of punctuated equilibrium is that A anagenesis can occur without cladogenesis 2 A photoheterotroph is an organism that obtains its energy A from sunlight and carbon from an inorganic source 3 All animals have at least one cluster of homeotic genes A 4 All species are defined as interbreeding groups of individuals A reproductively isolated from all others 5 Coenocytic hyphae lack internal divisions among cells A 6 Derived traits are usually more complex than ancestral traits A 7 Evolutionary convergence in 1 or more characters can lead to A polyphyletic groups in a phylogeny 8 Grades are acceptable groupings in a cladistic analysis A 9 Gram-positive bacteria lack peptidoglycan in their cell walls A 10 Paedomorphic heterochronic mutations have been important in A generating new lineages of organisms 11 Induction is the phenomenon by which the fate of cells in an A embryo is determined by interaction with other cells 12 Large and small genomes differ mostly in the number of genes A 13 Methanogens are a type of Archaea A 14 Multicellular algae are a polyphyletic group A 15 Oomycetes are convergent with fungi because they have chitin A in their cell walls 16 The 2 types of slime molds are plasmodial and colonial A 17 The chemical evolution hypothesis suggests that the A biochemistry of early life was significantly different than what is observed today 18 The earth is approximately 45 billion years old. A 19 The most diverse group of Eubacteria are the proteobacteria A 20 Two organisms in the same family but different genera share a A more recent common ancestor than two organisms in the same class but different orders False B B B B B B B B B B B B B B B B B B B B Section II: Multiple Choice. Please select the best answer. (1 mark each) 21. Which of the following statements about prions is false? A: prions can occur in both an innocuous and a harmful morph B: they are proteins that may be involved in cell signalling C: prions cause fatal diseases in non-immunized individuals D: prions lack RNA Page 1 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ ancestor A B C 22. In the diagram above, the figure on the far left represents a modern human adult. Figures A – C represent 3 adult specimens of hypothetical descendents of modern humans. What type of heterochronic mutation does each represent? A: B: C: D: A = neoteny; B = progenesis; C = postdisplacement A = progenesis; B = neoteny; C = hypermorphosis A = peramorphosis; B = deceleration; C = pedomorphosis A = predisplacement; B = hypermorphosis; C = postdisplacement 23. Which of the following is likely to act earliest in development? A: B: C: D: homeotic genes apoptosis cytoplasmic segregation induction 24. Which of the following statements about morphogens is false? A: morphogen gradients form when chemical signals diffuse along an axis B: the concentration gradient of a morphogen controls gene expression C: bicoid and nanos are morphogens that act early in Drosophila development D: morphogens are only found in animals (eumetazoa) 25. Which of the following statements about stromatolites is not true? A: the earliest stromatolites appeared more than 3 billion years ago B: stromatolites thrive in hypersaline conditions C: some prokaryotes in stromatolites produce O2 D: photosynthetic organisms in stromatolites do not reduce CO2 Page 2 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ β1 log y β α1 descendent ancestor α log x 26. The figure above represents the relative growth of two structures (Y and x) in a hypothetical organism. This is an example of: A: B: C: D: predisplacement progenesis acceleration hypermorphosis 27. Which of the following statements concerning the development of the Drosophila embryo is false? A: segmentation genes specify the number of body segments B: the products of maternal effect genes activate the expression of segmentation genes C: segmentation genes specify the correct polarity of each segment D: segmentation genes include gap genes, pair-rule genes and homeotic genes 28. Which of the following statements is false? A: 2 genes are homologous if they are related by descent from a common ancestral DNA sequence B: orthologs are genes (occurring in related species) that evolved from a common ancestral gene C: paralogs usually retain their original function, while function usually differs among orthologs D: paralogs are genes related by duplication within a genome Page 3 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ 29. Consider a large population that has been subdivided into two smaller subpopulations by a physical barrier. Which of the following does not contribute to speciation? A: genetic drift B: gene flow C: reproductive isolating mechanisms D: heterochronic mutations 1 2 3 4 5 6 7 group I = 6, 7 group II = 5, 6, 7 group III = 1, 2, 3, 4, 5, 6, 7 30. The figure above represents the hypothetical phylogeny of 7 species. The dashed lines indicate “deep time” (at the base of the tree is a distant ancestor). Which of the following statements is true? A: Group I is monophyletic, group II is paraphyletic, group III is polyphyletic. B: Group I is a clade, group II is a grade, group III is a clade. C: Group I is paraphyletic, group II is monophyletic, group III is polyphyletic. D: Group I is a clade, group II is a clade, group III is a clade. 31. Which of the following statements about Archaea is false? A: Archaea may not be a monophyletic group B: Archaea have unusual lipids in their cell membranes C: Archaea are characterized by having peptidoglycan in their cell walls D: Archaea are more closely related to Eukarya than to Eubacteria Page 4 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ 32. All birds have feathers, and only birds have feathers. Which of the following statements is true? A: Feathers are ancestral when considering evolutionary relationships among birds, but derived when placing birds within a phylogeny including other groups B: Feathers are a derived trait used to determine how flightlessness evolved in some birds C: Feathers are a derived trait when comparing extinct birds to modern birds D: Feathers are an ancestral trait when considering the evolutionary relationships among all vertebrates 33. Which of the following statements about prokaryotic cells is false? A: Prokaryotes are permanently haploid B: Prokaryotes lack a cytoskeleton C: Prokaryotes lack membrane-bound organelles D: Prokaryotes undergo rapid mitosis 34. Which of the following statements about hsp90 is false? A: hsp90 is a chaperone protein B: hsp90 may improve the function of some proteins C: hsp90 encodes a transcription factor D: hsp90 keeps some proteins unstable 35. Consider a lineage of marine snails. During evolution, there were a number of speciation events in this lineage, resulting in variation in body size among the daughter species. The larger snails were often more successful than the smaller snails, resulting in frequent adaptive radiations among the larger snails. This is an example of: A: differential extinction B: punctuated equilibrium C: anagenesis without cladogenesis D: differential speciation 36. Which of the following types of characters are useful in a cladistic analysis? A: autapomorphies and synapomorphies B: shared derived characters C: all types of characters (morphological, biochemical, etc.) D: ancestral and derived traits Page 5 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ Section III. Match the group on the left with the characteristic on the right. Each letter may be used once, more than once, or not at all. Each question is worth ½ mark. Q 37 38 39 40 41 Organism: firmicutes C spirochetes D purple bacteria A cyanobacteria B Chlamydia E Characteristic A: non-oxygenic photosynthesis B: oxygenic photosynthesis C: endospores D: axial filaments E: obligate intracellular parasites Q: 42 43 44 45 46 Organism: diplomonads B euglenozoans A dinoflagellates D ciliates E diatoms C Characteristic A: crystalline rod B: multiple flagella C: have photosynthetic endosymbionts D: responsible for “red tide” E: contractile vacuoles Q: 47 48 49 50 51 Organism: Chytridiomycota B Zygomycota C Glomeromycetes D Ascomycota E Basidiomycota A Q: 52 53 54 55 56 Term: apoptosis B Ubx A induction D cytoplasmic segregation C aniridia E Characteristic A: bracket fungi and most mushrooms B: motile spores C: include the microsporidia D: all form mycorrhizae E: symbiosis with algae, cyanobacteria Example A: mutation results in development of extra wings B: failure results in syndactyly C: controls location of cleavage D: responsible for formation of optic cup E: homolog of Pax-6 and eyeless Page 6 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ Section IV. Short answer questions. Choose 1 of the following 5-mark questions. Please answer in the space provided (remember, longer is not always better!). 57. Consider the following matrix for 10 characters of 4 species (A – D). 0 = absence and 1 = presence of a particular trait. Construct a phenogram based on these data. (Show your rough work.) 1 2 3 4 5 6 7 8 9 10 A B 0 0 1 0 0 0 0 1 0 0 0 0 1 1 1 0 0 0 1 1 C 1 1 1 0 1 1 0 1 1 0 D 1 1 1 1 1 1 0 1 0 0 Constructing Taxon Matrix (2 marks) A B C D • • • A - B 0.7 - C 0.3 0.0 - D 0.3 0.2 0.8 - -1 mark for 1st mistake -2 marks for >2 mistakes -1 mark for not converting to decimal or fraction Joining Closest Groups C-D & A-B (2 marks) (A-D)+(B-D)+(A-C)+(B-C) 4 • • (0.3)+(0.2)+(0.3)+(0.0) = 0.8 = 0.2 4 4 -1 for including equation -1 for incorrect solution Phenogram (1 mark) A B C D Page 7 of 8 Name:_________________________________ Student #__________________ Lab Section:_____________________________ TA: ______________________ 58. What were the advantages of the 5 kingdom system of classification over previous classification systems? Why is it now considered obsolete? A) 5 kingdom system: 2 domains (Prokaryotes and Eukaryotes); one kingdom of prokaryotes (Monera) and 4 eukaryotes (Protoctista, Fungi, Plantae and Animalia) • 0.25 marks per point • no marks for ‘5 kingdom system’ without reference to other points B) Advantages: recognized fundamental split between pro- and eukaryotes; saw “protoctists” as basal to other eukaryotes; recognized differences in nutritional modes among eukaryotes; Fungi separate from Plantae; “protozoans” removed from Animalia; algae removed from Plantae • 0.25 marks per point C) Problems: lacked cladistic approach: monera & protoctists were paraphyletic groups • 0.5 marks for cladistics point • 0.5 marks for example (monera & protistis) D) Obsolete because differences within “Monera” more profound than between some “monerans” and eukaryotes • 1 mark E) New approach recognizes 3 domains (Archaea, Eubacteria and Eukarya) • 0.5 marks for mentioning 3 domains • 0.5 marks for naming them 59. Describe the hypothesized steps that occurred in the evolution of the eukaryotic cell. A) Loss of cell wall • 1 mark B) Infolding of cell membrane (perhaps enclosing “nucleus”, ER, golgi apparatus, lysosomes) • 1 mark • -0.5 for not mentioning ‘cell membrane’ C) Formation of flagella, cilia, etc. • 1 mark D) Cytoskeleton (actin + microtubules) • 1 mark E) Endocytosis of peroxisomes, mitochrondria, chloroplasts • 1 mark • -0.5 marks for not mentioning peroxisomes, mitochrondria or chloroplasts) Page 8 of 8