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Life’s Diversity through Evolution Science vs. Religion What is science based on? Science is based on OBSERVABLE evidence. What is religion based on? Religion is based on FAITH. – By definition, faith is something that is not observable. Since they are 2 different “fields,” they are not competing with each other Religious Scientists? My religion consists of a humble admiration of the illimitable superior spirit who reveals himself in the slight details we are able to perceive with our frail and feeble mind. – Albert Einstein (1879 - 1955) Science without religion is lame, religion without science is blind. – Albert Einstein (1879 - 1955), "Science, Philosophy and Religion: a Symposium", 1941 Not only does God play dice, but... he sometimes throws them where they cannot be seen. – Stephen Hawking Evolution is a Theory Theory Defined: “scientific principle to explain phenomena: a set of facts, propositions, or principles analyzed in their relation to one another and used, especially in science, to explain phenomena” – MSN Encarta, http://encarta.msn.com/dictionary_/theory.html Evolution is as much a theory as… – Cell theory: biology – Atomic theory: chemistry – Plate tectonic theory: geology A Final Quote “Be opinionated, just not ignorant.” Evolution The theory that organisms on Earth have changed over time is known as evolution. Charles Darwin (1809-1882) is the man who contributed more to our understanding of evolution than any other individual. Charles Darwin English naturalist (scientist) 18091882 1831, Darwin took a job as a naturalist on the HMS Beagle – Went on a 5-year scientific journey around the world He collected many biological and fossil specimens – Combined with his previous and subsequent observations Published On the Origin of Species by Natural Selection in 1859 – These ideas are a basic unifying theme of biology today – Without evolution, biology doesn’t make as much sense He realized that he had never seen many of these organisms and the diversity of life on Earth was enormous! Darwin questioned “Where did all of these different life forms come from?” EX: Darwin collected 68 species of beetles from Brazil in just a few hours while on shore. Darwin also discovered many fossils of organisms that no longer existed today. This lead to other questions, “Where did all these organisms go? And why did they disappear?” Fitness Darwin observed that many plants and animals had parts and displayed behaviors that helped them survive. Traits and behaviors that will help organisms survive and reproduce in their environment give those organisms greater fitness (ability to survive and reproduce). Those organisms that are most fit will survive and reproduce more often than those organisms who are “less” fit. Became known as “survival of the fittest.” – Darwin did not use that phrase, it came about later & may lead to misconceptions… Fitness Darwin argued that increased fitness arises from adaptations. Certain adaptations allow organisms to become better suited to their environment, and thus better able to survive and reproduce. Adaptations can be physical or behavioral Does “struggle for existence” mean only the biggest, strongest animals survive? What types of characteristics increases an organisms chance of surviving? Adaptations Adaptation – Evolution of a structure, behavior, or internal process that enables an organism to respond to environmental factors and live to produce offspring – Can also be the structure itself Structural adaptations take a long time Physiological adaptations can happen faster – Bacteria strains develop resistances to antibiotics fairly quickly Summary of Darwin’s Idea There is variation between individuals. Summary of Darwin’s Idea Organisms tend to have more offspring than can possibly survive, so there is competition among the offspring for a limited amount of resources Summary of Darwin’s Idea Organisms compete for limited resources There is a struggle for existence Summary of Darwin’s Idea Those organisms that are best suited for their environment (most fit) will survive and pass on their genes. Darwin’s Theory of Evolution Darwin developed evolution by natural selection Compared the process of natural selection in nature vs. artificial selection used by people. He used this comparison to develop a hypothesis to explain how evolution occurs. Using artificial selection a breeder can produce plants and animals that look very different from their ancestors. Artificial selection uses the variation nature provides. How does horse racing relate to selective breeding? Darwin’s Origin of Species He filled notebooks with ideas about species diversity & the process of evolution. His beliefs challenged the fundamental ideas of that time (species do not change). Darwin was very reluctant to publish his book due to his fear of what the public would think. Are their any scientific topics today that are controversial? Origin of Species 1858, a naturalist name Alfred Wallace sent him a letter describing evolution. He had been working in Malaysia and come to the same conclusions Darwin did. Darwin had been working on his theory and the evidence to support it for 20 years. Darwin published the Origin of Species in 1859, 23 years after his trip. Origin of Species His book did two things: 1. Proposed a mechanism for evolution (natural selection) 2. Evidence showing it had been taking place for millions of years and continues to take place Descent with Modifications Over long periods of time, natural selection will start to produce organisms with different structures or organisms that occupy different niches or habitats. Because of this, species today look different from their ancestors. Selection Artificial selection – Breeding organisms with specific traits in order to produce offspring with identical traits Natural selection – A mechanism for change in populations – Occurs when organisms with favorable traits survive, reproduce and pass their traits on to the next generation Organisms with these traits are said to be more “fit” Organisms without these traits are less likely to survive and Darwin awards Geological Evolution – James Hutton (1788) proposed that rain, wind, temperature, and natural disasters shaped the valleys, mountains, and rocks over extremely long periods of time – Charles Lyell (1830)contributed to this by adding that current observations of volcanic eruptions and other natural disasters shows that Hutton was correct in assuming the Earth was older than originally predicted Acceptance of Geological Evolution Important to Darwin because it established an OLD Earth Darwin’s theory of evolution required lots of time Another important influence: Thomas Malthus – Observed that humans were being born at a faster rate than they were dying – War, famine, and disease control populations – These observations provided enough curiosity for Darwin to delve deep into his theory of Natural Selection humans evolve First placental mammals Flowering plants dominant First birds 1st dinosaurs First reptiles First seed plants First amphibians First vertebrates First invertebrates eukaryotes prokaryotes Life evolves Dating Techniques Geologic Record – Hypothetical library of all the known geological processes on Earth – Determined through inference and dating methods Relative Dating (hee hee) – Sediments are constantly being laid down. – Over time, many layers form – If a fossil is in a layer below a different fossil, it is older than that fossil Radiometric Dating Radiometric Dating – Radioactive isotopes decay and form new isotopes - the rate this happens is called the isotope’s half life – Ex: Potassium-40 decays to argon-40 and has a half life of 1.3 billion years Half a sample will decay to argon-40 in 1.3 billion years So if there are equal amounts of potassium-40 and argon-40 in an area, the sample has been in place 1.3 billion years – Carbon 14 has a half life of only 7,000 years for more recent sample dating Review 1. 2. 3. 4. How does the geologic record help us? What use is relative dating? What is radiometric dating? Why are different isotopes used in radiometric dating? Evidence for Evolution Fossil Record – A hypothetical “library” of all the fossils collected around the world Fossils show changes throughout time – 99 percent of all animals are now extinct – Fossil record shows ancestors with similar characteristics Embryology Study of embryos and the relationships that exist between different species Evidence for Evolution: Embryology Many species share features in the young embryos – Ex: Mammals, reptiles, birds and fish all have a tail and pharyngeal pouches (develop into different respiratory systems) Evidence For Evolution: Anatomy Homologous Structures are structural features with a common evolutionary origin – Ex: whale forelimb, crocodile forelimb, bird wing, human forelimb all look the same – Can you think of other examples? Analogous Structures are body parts or organisms that do not have a common evolutionary origin but are similar in function – Bird, bat and butterfly wings – Show how organisms adapt to different ways of life and different conditions Homologous Structures Structure and Function between species is the same Analogous Structures Functions are the same between species; structures are different Evidence for Evolution Vestigial Structures are body structures that no longer serve their original purpose, but was probably useful to an ancestor – Ex: human appendix, pelvic bone in baleen whale, “tail” in humans, some human’s ability to wiggle their ears Vestigial Structures Evidence shows the organ was once used, but not any longer Evidence for Evolution Biochemistry Nearly all organisms share DNA, ATP, and many enzymes among their biochemical molecules Organisms that share more similar enzymes, DNA sequences, etc. are more closely related Review 1. 2. 3. 4. 5. 6. How did Darwin come up with his ides for natural selection? Some snakes have vestigial legs. Why is this considered evidence for evolution? Explain how adaptations such as camouflage help species survive. How do homologous structures provide evidence for evolution? Why did birds and bats both get wings? A parasite that lives in red blood cells causes the disease called malaria. In recent years, new strains of the parasite have appeared that are resistant to the drugs used to treat the disease. Explain how this could be an example of natural selection occurring. Population Genetics Populations evolve, not individuals Natural selection acts on a range of phenotypes in a population Evolution is often defined as a change in the frequency of an allele in a population over time Population Genetics All of the population’s genes are in a theoretical “gene pool” The percentage a specific allele in the gene pool is called the allelic frequency A population in which the frequency of alleles remains relatively the same over generations is in genetic equilibrium Any change in this genetic equilibrium results in evolution Hardy-Weinberg Principle 1. 2. 3. 4. 5. How to Stop Evolution – must meet all five requirements Population is so large, chance alone cannot change relative frequency (genetic drift doesn’t occur) Mutations do not occur All genotypes have equal fitness (no natural selection) No organisms leave or enter a population Mating occurs at random Mutations Mutation provides the raw material for evolution to act upon – Poor mutations are selected against, good mutations are selected for – Change the frequency of alleles – Examples: Genetic Drift Genetic drift is the alteration of allelic frequencies by chance events Genetic drift can greatly affect small populations – Ex: The Amish community carries an allele that results in short arms and legs and extra fingers and toes (1/14 vs. 1/1000) Natural Selection Still the most significant factor that causes changes in established gene pools Stabilizing Selection – Natural selection that favors average individuals (on a normal curve) in a population – Reduces variation in a population – Ex: With spiders, larger ones are found easier and eaten and small spiders can’t find food as easily Natural Selection Directional Selection – Natural selection favors one of the extreme variations of a trait – Can lead to rapid evolution – Ex: The food supply in an area is limited to hard nuts. Birds with short, strong beaks will survive Disruptive Selection – Natural selection favors individuals with either extreme of a trait’s variation – Can lead to evolution of 2 new species – Ex: A shelled, marine organism called a limpet has white, tan, and dark shells. The white and dark shells blend in on different colored rocks. Tan gets eaten. Species A species is a group of organisms that look alike and can interbreed to produce fertile offspring in nature Speciation is the evolution of new species – Occurs when members of similar populations no longer interbreed to produce fertile offspring within their natural environment Causes of Speciation Physical barriers – Volcanic eruptions, sea-level changes, new islands forming These prevent interbreeding Called geographic isolation Over time, may have to adapt to new types of environments Causes of Speciation Reproductive Isolation Two types – One occurs because of geographical reasons (migrate away) – One occurs because of behavioral reasons Some mate in fall, some in spring Polyploidy Individual with a multiple of a normal set of chromosomes How does this happen? New zygotes may not develop the same way as parents due to different number of chromosomes May result in speciation Gradualism James Hutton & Charles Lyell Theory that processes are moving at the same speed today as in the past Suggests that the Earth is very old – 4.55 billion years Also suggests that evolution occurs constantly, but slowly Punctuated Equilibrium Remember first day of class? Niles Eldredge and Steven J. Gould Theory that speciation occurs relatively, in rapid bursts, with long periods of genetic equilibrium inbetween Might occur due to drastic environmental changes resulting in quick adaptations Adaptive Radiation When an ancestral species evolves into an array of species to fit a number of diverse habitats Hawaiian Island honeycreepers – Similar in body size and shape, but different sharply in color and beak shape – Adapted to occupy different niches (what’s a niche) Divergent Evolution Adaptive radiation is a type of divergent evolution – The pattern of evolution in which species that once were similar to an ancestral species diverge, or become increasingly distinct Galapagos Finches – Similar changes for finches that ended up on different small islands Convergent Evolution A pattern of evolution in which distantly related organisms evolve similar traits Occurs when different organisms occupy similar environments – Adapt similar traits Example: – Organ pipe cactus in N. and S. American vs. Euphorbiaceae in African deserts both look very similar Review 1. 2. 3. 4. 5. 6. Explain and illustrate why the evolution of resistance to antibiotics in bacteria is an example of directional natural selection. How can geographic isolation change a population’s gene pool? Why is rapid evolutionary change more likely to occur in small populations? How do gradualism and punctuated equilibrium differ? How are they similar? Hummingbird moths are night-flying insects whose behavior and appearance are similar to those of hummingbirds. Explain how these two organisms demonstrate convergent evolution. What is divergent evolution? How does it compare to adaptive radiation? Quick Tour of Ch 17 Classification – the grouping of objects or information based on similarities Taxonomy – the branch of biology that groups and names organism based on studies of their different characteristics Aristotle was the first on record – Classified organisms into two groups: plants and animals, each with subgroups Carolus Linnaeus Developed taxonomic system still used today Based on physical and structural similarities of organisms Grouped life into 7 taxa (groups) – What are they? Kingdom, phylum, class, order, family, genus, species Five Kingdoms: animals, plants, fungi, protists, monerans A six Kingdom system is becoming more and more common: monera split into eubacteria and archaebacteria Binomial Nomenclature All organisms have a two-word name Consists of their genus and their species – Always italicized in type and underlined in writing Genus is capitalized, species is lower case – Homo sapiens (human) – Passer domesticus (sparrow) Classification Key Sometimes called a dichotomous key “Guide” for identifying different species based on observable traits Useful for distinguishing between types of organisms Modern Classification Generally based on evolutionary relationships vs. physical characteristics Used structural similarities, breeding behavior, geographical distribution, chromosome comparisons, biochemistry to figure out relationships Phylogenetic Diagram Monerans evolved first (often broken up into archaebacteria and eubacteria) – prokaryotes Protists then evolved – Single celled eukaryotes – Three general kinds of protists: fungus like, animal like and plant like Fungi, Plants and Animals evolved from the types of protists Review 1. 2. 3. 4. 5. Why does your book use the classification system of six kingdoms instead of five? What two taxa does binomial nomenclature consist of? Why is classification hard to do? What are the 7 Linnaean taxa? How might you use taxonomic key to figure out the name of an organism?