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Chapter 10 Principles of Evolution Early scientists proposed ideas about evolution. • Evolution is the biological change process by which descendants come to differ from their ancestors. • A species is a group of organisms that can reproduce and have fertile offspring. • There were many important naturalists in the 18th century. – Linnaeus: classification system from kingdom to species – Buffon: species shared ancestors rather than arising separately – E. Darwin: more-complex forms developed from lesscomplex forms – Lamarck: environmental change leads to use or disuse of a structure Theories of geologic change set the stage for Darwin’s theory. • The age of the Earth was a key issue in the early debates over evolution • Common view was that Earth was created about 6000 years ago and the Earth nor its species have changed • There were three theories of geologic change. – catastrophism – gradualism – uniformitarianism • The theory of catastrophism states that natural disasters such as floods and volcanic eruptions have happened often during Earth’s history. • Shaped landforms and caused species to become extinct • James Hutton proposed that changes in landforms were the result of slow changes over a long period of time – principal of gradualism • Uniformitarianism is the prevailing theory of geologic change.(Lyell) • The geologic processes that shape Erath are uniform through time Darwin’s observations Charles Darwin the Naturalist 10 Charles Darwin and his Voyage on the Beagle • Born Feb. 12, 1809 • Joined Crew of HMS Beagle, 1831 • Naturalist • 5 Year Voyage around world • Avid Collector of Flora & Fauna • Astounded By Variety of Life 11 A reconstruction of the HMS Beagle sailing off Patagonia. 12 Darwin Left England in 1831 Darwin returned 5 years later in 1836 13 HMS Beagle’s Voyage 14 The Galapagos Islands • Volcanic islands off the coast of South America • Very different climates • Island species varied from mainland species & from island-to-island species • Animals On Islands Unique • Tortoises • Iguanas • Finches 15 Darwin observed differences among island species. • Variation is a difference in a physical trait. • Galápagos tortoises that live in areas with tall plants have long necks and legs. • Galápagos finches that live in areas with hard-shelled nuts have strong beaks. 17 The Finches • Finches on the islands resembled a mainland finch • More types of finches appeared on the islands where the available food was different (seeds, nuts, berries, insects…) • Finches had different types of beaks adapted to their type of food gathering 18 19 • An adaptation is a feature that allow an organism to better survive in its environment. – Species are able to adapt to their environment. – Adaptations can lead to genetic change in a population. Darwin observed fossil and geologic evidence supporting an ancient Earth. • Darwin found fossils of extinct animals that resemble modern animals. • Darwin found fossil shells high up in the Andes mountains. • He saw land move from underwater to above sea level due to an earthquake. • Darwin extended his observations to the evolution of organisms. Theory of Natural Selection KEY CONCEPT Darwin proposed natural selection as a mechanism for evolution. Several key insights led to Darwin’s idea for natural selection. • Darwin noticed a lot of variation in domesticated plants and animals. • Artificial selection is the process by which humans select traits through breeding. neck feathers crop tail feathers • Natural selection is a mechanism by which individuals that have inherited beneficial adaptations produce more offspring on average than do other individuals. • Heritability is the ability of a trait to be passed down. • There is a struggle for survival due to overpopulation and limited resources. • Darwin proposed that adaptations arose over many generations. Natural selection explains how evolution can occur. • There are four main principles to the theory of natural selection. – variation – overproduction – adaptation – descent with modification • Fitness is the measure of survival ability and ability to produce more offspring. ADAPTATION VARIATION OVERPRODUCTION DESCENT with MODIFICATION Natural selection acts on existing variation. • Natural selection can act only on traits that already exist. • Structures take on new functions in addition to their original function. five digits wrist bone Evidence for Evolution KEY CONCEPT Evidence of common ancestry among species comes from many sources. Evidence for evolution in Darwin’s time came from several sources. • Fossils provide evidence of evolution. • Fossils in older layers are more primitive than those in the upper layers. • The study of geography provides evidence of evolution. – island species most closely resemble nearest mainland species – populations can show variation from one island to another • Embryology provides evidence of evolution. – identical larvae, different adult body forms – similar embryos, diverse organisms Larva Adult crab Adult barnacle • The study of anatomy provides evidence of evolution. – Homologous structures are similar in structure but different in function. – Homologous structures are evidence of a common ancestor. Human hand Mole foot Bat wing • The study of anatomy provides evidence of evolution. – Analogous structures have a similar function. Human hand foot • Analogous structures are notMole evidence of a common ancestor. Fly wing Bat wing Structural patterns are clues to the history of• aVestigial species. structures are remnants of organs or structures that had a function in an early ancestor. • Ostrich wings are examples of vestigial structures. Evolutionary Biology Today Fossils provide a record of evolution. • Paleontology is the study of fossils or extinct organisms. • Paleontology provides evidence to support evolution. Molecular and genetic evidence support fossil and anatomical evidence. • Two closely-related organisms will have similar DNA sequences. • Pseudogenes are sequences providing evidence of evolution. – no longer function – carried along with functional DNA – can be clues to a common ancestor • Hox genes indicate a very distant common ancestor. – control the development of specific structures – found in many organisms • Protein comparisons, or molecular fingerprinting reveals similarities among cell types of different organisms. Evolution unites all fields of biology. • Scientist from any fields contribute to the understanding of evolution. • The basic principles of evolution are used in many scientific fields.