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Population Genetics and Speciation 02.05.08 / 02.06.08 Genetic Equilibrium Population biologists study many different traits in populations, such as size and color Causes of variation: Traits vary and can be mapped along a bell curve, which shows that most individuals have average traits, whereas a few individuals have extreme traits. Variations in genotype arise by mutation, recombination, and the random pairing of gametes. Bell Curve Genetic Equilibrium The total genetic information available in a population is called the gene pool. Allele – one of the alternative forms of a gene that governs a certain characteristic Allele frequency is determined by dividing the total number of a certain allele by the total number of alleles of all types in the population. Genetic Equilibrium Phenotype – a detectable characteristic that results from genotype and environment Phenotype frequency is equal to the number of individuals with a particular phenotype divided by the total number of individuals in the population. Genetic Equilibrium Hardy-Weinberg genetic equilibrium is a theoretical model of a population in which no evolution occurs and the gene pool of the population is stable. Serves as the null hypothesis Allele frequencies in the gene pool do not change unless acted upon by certain forces. Genetic Equilibrium Phenotypes (red, pink, white) and alleles (R and r) Disruption of Genetic Equilibrium Evolution may take place when populations are subject to genetic mutations, gene flow, genetic drift, nonrandom mating, or natural selection. Mutations are changes in the DNA. There are an estimated 1.6 allele mutations created with each person Disruption of Genetic Equilibrium Gene Flow is the process of genes moving from one population to another Emigration and immigration cause gene flow between populations and can thus affect gene frequencies. Genetic drift is a change in allele frequencies due to random events. operates most strongly in small populations. Disruption of Genetic Equilibrium Nonrandom mating Mating is nonrandom whenever individuals may choose partners. There are two types of selection: sexual selection natural selection Disruption of Genetic Equilibrium Sexual selection occurs when certain traits increase an individual’s success at mating. Sexual selection explains the development of traits that improve reproductive success but that may harm the individual. Disruption of Genetic Equilibrium Natural selection can influence evolution in one of three general patterns. Stabilizing selection favors the formation of average traits. Disruptive selection favors extreme traits rather than average traits. Directional selection favors the formation of more-extreme traits. Disruption of Genetic Equilibrium Graphic Organizer Populations can evolve through gene mutation gene flow nonrandom mating genetic drift natural selection Formation of Species According to the biological species concept, a species is a population of organisms that can successfully interbreed but cannot breed with other groups. Donkeys and horses can produce mules, but mules are sterile Fitness is measured in grandchildren Formation of Species Geographic isolation results from the separation of population subgroups by geographic barriers. Geographic isolation may lead to allopatric speciation. allo (different) and patric (patriot) Reproductive isolation results from the separation of population subgroups by barriers to successful breeding. Reproductive isolation within the same geographic area is known as sympatric speciation. sym (together) and patric (patriot) Formation of Species In the gradual model of speciation (gradualism), species undergo small changes at a constant rate. Under punctuated equilibrium, new species arise abruptly, differ greatly from their ancestors, and then change little over long periods.