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How Evolution Works Genes, Geography and Sex Early Idea: Lamarckian Evolution Fossil record indicated increasing complexity Reasoned traits are passed via use and disuse e.g. the giraffe’s neck Step 1: Raw Material Genes are raw material Forms of genes = alleles Polygenic vs. Single gene trait Selection on a Single Gene: The Peppered Moth Before Industrial Revolution After Industrial Revolution Single gene controlling a trait will have only a few (usually 2 or 3 phenotypes) Variation and Selection Variation from two sources 1) New mutations = new allele types 2) Gene shuffling = new allele combinations Any change in allele frequency = Evolution Peppered Moth Simulation Polygenic Traits More than one gene controls a trait Selection and Changing the Norm Most traits are polygenic The normal trait is the average or mean in the population Selection changes the mean, usually lowers variation Selection will adjust mean Stabilizing Selection Disruptive Selection Can lead to new species Does Evolution Ever Stop? YES, but only if the following conditions are met 1) Random mating 2) Large Population 3) No movement in or out of population 4) No Mutations 5) No Natural Selection Hardy Weinberg Equilibrium How New Species Are Formed Two populations must become reproductively isolated Behavioral Isolation Groups differ in mating, feeding, sleep/awake cycles so members of populations do not meet to mate Rana aurora (Redlegged frog) Breeds in fast moving streams Rana catesbiana (Common bullfrog) Breeds in still ponds Geographic Isolation Geological Time and Evolution First fossils 3.5 bya Stromalites 550 mya = Cambrian Explosion (by fossil evidence) Dating Fossil Age Relative Dating Radioisotope Dating Known decay times of isotopes can be used Extinction and Adaptive Radiation Most species cannot adapt Those that can radiate into open niches Evolution Patterns Convergent Evolution Coevolution