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UNIT 3: Evolution Chapter 7: Introducing Evolution Chapter 8: Developing a Theory of Evolution What is the theory of evolution, and what evidence is there to support it? Chapter 9: Evolution and Speciation UNIT 3 Chapter 8: Developing a Theory of Evolution 8: Developing a Theory of Evolution Any scientific theory requires evidence to support its assumptions or postulates. Fossils, such as these trilobites, support the scientific theory of evolution as it relates to life on Earth. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 8.1 Scientific Contributions to a Theory of Evolution John Ray (1627-1705) • one of the first scientists to carry out empirical studies on the natural world • developed a classification system for plants and animals based on anatomy and physiology (refined by Linnaeus) Georges-Louis Leclerc, Comte de Buffon (1707-1788) • one of the first people to publicly challenge the idea that life forms are unchanging • speculated that Earth was more than 6000 years old UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Scientific Contributions to a Theory of Evolution Mary Anning (1799-1847) • worked as a fossil hunter and uncovered the first plesiosaur • work was confirmed by Georges Cuvier, the founder of paleontology Mary Anning was very skilled at collecting fossils. As a result, she made many important contributions to the field of paleontology. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Catastrophism Georges Cuvier (1769-1832) • credited with founding the field of paleontology (study of fossils) • discovered that each stratum (layer) of rock held a unique group of fossil species • discovered that the oldest fossils are in the deepest layer • suggested that catastrophes killed many species (catastrophism) and that these events corresponded to the boundaries between the fossil strata Deep rock strata are older than strata closer to the surface. Different species of fossilized organisms can be found in different sedimentary rock strata. This is evidence that not all life forms came into existence at the same time. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Uniformitarianism Charles Lyell (1797-1875) • rejected Cuvier’s theory of catastrophism • proposed the theory of uniformitarianism: geological processes in the past operate at the same rate as they do today • suggested that a slow and continuous process could result in substantial changes in the long term • agreed that Earth was more than 6000 years old UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Line of Descent Jean-Baptiste Lamarck (1744-1829) • proposed a “line of descent” progression between current species and fossil forms • suggested species increased in complexity and became better adapted to their environment over time • proposed the idea of the inheritance of acquired characteristics whereby characteristics acquired during an organism’s lifetime could be passed on to offspring UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Evolution by Natural Selection Thomas Malthus (1766-1834) • an economist who wrote Essay on the Principles of Population that explained how populations might change over time • key idea was that populations produce far more offspring than their environments can support Charles Darwin (1809-1882) • travelled the coast of South America and made natural and geographical observations • used these observations to propose the theory of natural selection: how life has changed and continues to change, due to natural pressures, the first comprehensive presentation of the idea Alfred Russel Wallace (1823-1913) • made observations and conclusions similar to Darwin’s • Darwin and Wallace recognized the importance of Malthus’s essay UNIT 3 Chapter 8: Developing a Theory of Evolution Evolution by Natural Selection The five-year voyage of the HMS Beagle took Darwin around much of the world. He spent most of his time exploring the coast and coastal islands of South America. Section 8.1 UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Evolution by Natural Selection The theory of evolution by natural selection includes the idea of survival of the fittest as discussed in Darwin’s The Origin of Species: • Organisms produce more offspring than can survive. Therefore, organisms compete for limited resources. • Individuals of a population vary extensively, and much of this variation is heritable. • Individuals that are better suited (fitted) to local conditions survive to produce more offspring. • Processes for change are gradual. The changes in populations can be viewed as descent with modification, as the change does not demonstrate progress (improvement). It is simply change. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Darwin’s Observations and Questions Some of Darwin’s observations that led to his theory were: • The flora and fauna were different in different regions. • He found fossils of extinct animals that looked similar to living animals. • Finches found on islands resembled continental finches but were different in some characteristics. • Finch beaks were adapted to the food source on each island. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Darwin’s Observations and Questions Questions that Darwin considered: • Why were all types of organisms not randomly distributed? • Why would living and fossilized organisms that looked similar be found in the same region? • Why did the Galapagos species so closely resemble organisms on the adjacent South American coastline? • Could species have been modified from an ancestral form that arrived on the Galapagos Islands shortly after the islands were formed? UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.1 Review Section 8.1 UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 8.2 Sources of Evidence for Evolution The fossil record reveals the history of life on Earth. Continued… UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Sources of Evidence for Evolution UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Fossil Evidence The fossil record provides the following evidence: • Fossils in younger (and usually closer to the surface) rock are more similar to current species than those in deeper strata. • Fossils appear in “chronological order” in the rock strata, and probable ancestors of a species are found in older rocks. • Not all organisms appear in the fossil record at the same time, showing the successive evolution of groups (kingdoms) of species. Paleontologists have used fossils to trace the evolution of the modern camel. The Paleocene, Eocene, Oligocene, and Miocene epochs are subdivisions of the Cenozoic era. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Fossil Evidence Scientists are always looking for transitional fossils that show an intermediary link between past and present groups of organisms. Vestigial structures are reduced versions of what were once functional structures in an ancestral species. A key piece of evidence is a fossil record that shows the once functional use of a present day vestigial structure. Fossil evidence suggests that modern toothed whales evolved from a terrestrial ancestor, Pakicetus attocki. Basilosaurus and Dorudon, not shown in this illustration, appear more recently in the fossil record, after the appearance of Rodhocetus. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Evidence from Biogeography Biogeography is the study of the past and present geographical distribution of organisms. Darwin and Wallace supported their theories with biogeographical evidence showing that species evolved in one location and spread out to other regions. • Geographically close environments are more likely to be populated by related species. • Animals found on islands often closely resemble animals on the nearest continent. • Fossils of the same species can be found on the coastlines of neighbouring continents. As the southern supercontinent Gondwana broke apart about 150 million years ago, the land masses that became the current continents of Africa, Australia, South America, and Antarctica were isolated from each other. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Evidence from Anatomy The study of anatomy provides the following evidence: • Homologous structures are those that have similar structural elements and origin but may have a different function. They originate from a common ancestor. Examples include forelimbs in vertebrates and mammalian hairs. • Analogous structures do not have a common evolutionary origin but perform similar functions. These provide evidence for adaptation to suit the environment. An example is the fins of a porpoise and a fish. What type of anatomical evidence of evolution is shown in the images? UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Evidence from Embryology Embryology is the study of pre-birth stages of an organism’s development. It has been used to determine evolutionary relationships between animals: • Generally, related species share common embryological stages and timings of structural development. • All vertebrate embryos have paired pouches, or outpocketings, of the throat (these later develop into ears/throat or gills). Similarities in the embryos of fish, birds, and mammals provide evidence of evolution of species from a common ancestor. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Evidence from DNA (Molecular Evidence) Evolutionary relationships are reflected in DNA since DNA carries genetic information from generation to generation: • Species that have similar DNA share a common ancestor. • DNA evidence supports conclusions about relationships and common ancestry provided by other areas of evidence. UNIT 3 Chapter 8: Developing a Theory of Evolution Section 8.2 Review Section 8.2 UNIT 3 STSE Feature: Biology Connections T. rex and chickens share a common ancestor?