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Theory of Evolution by Natural Selection NATURAL SELECTION is the way in which nature favours the reproductive success of some individuals within a population over others PROCESS OF NATURAL SELECTION (variation , selection, inheritance, time) 1. VARIATION WITHIN POPULATIONS: Individuals within a population can contain many different traits/adaptations. Genetic variation occurs by chance through genetic mutations and recombination. 2. SELECTION: Through competition or change the environment can act upon individuals differently depending on the traits they posses. Healthy populations produce far more offspring than could survive and reproduce. 3. INHERITANCE: parents pass on their traits to their offspring genetically. As a result of selective pressures, some individuals would have a better chance of survival 4. TIME: Since traits can be heritable, those that survive live long enough can reproduce and pass on their traits. Over time the POPULATION CHANGES. The more advantageous trait, which allows for survival and more offspring, will become more common in the population. If this process continues, eventually, most individuals in the population will have the advantageous trait Darwin’s Observations: Finches An ancestral population of medium-size billed ground finches are amongst the first seed eating birds to arrive on the Galapagos Islands. These birds had access to all types of seeds ( mostly large and some medium) and had little to no competition Over time, the finch population grew to it maximum carrying capacity. New born finches born with variability (some slightly lager bills and some slightly smaller than average sized bills) were exposed to competition within the species Most of the birds had to compete for the medium sized seeds, while those with the larger bills were able to utilize the larger seeds as a food source Generation after generation, the average bill size continues to increase. MODERN TAKES ON THE THEORY Survival of the fittest is a term used to describe the process of natural selection (Herbert Spencer, a contemporary of Darwin coined this term). An individual that is considered fit, has adaptations that allow it to survive longer and favour reproduce success. Adaptation is a result of a characteristic or feature of a species that makes it well suited for survival or reproductive success in its environment Examples: A white coat of a polar bear provide an adaptation for camouflage The long snout of an anteater provides it with an adaptation to better access its food source The thick blubber of a seal provides it with an adaptation to survive colder temperatures of its habitat The size and shape of a ground-finch’s beak determines which food sources it can access and provide an adaptation that makes them suitable for specific niches ( a habitat suitable for its behaviour and food preferences) TYPES OF SELECTION: Directional selection: o Selection that occurs when selection favours individuals with a more EXTREME variation of a trait. The result is a shift in the average condition o Example: The beak size of the ground finches was a result of directional selection toward the larger beak size. o Example: Hummingbirds exposed to a new habitat with longer flowers. Individuals with slightly longer bills will be more successful or “fit” in comparison to those with small or medium bills as the environmental pressures favour long bills. Stabilizing selection: o Occurs when the AVERAGE phenotype within a population is favoured by the environment. Selection against individuals exhibiting traits that deviate from the current population average occurs. o Example: Hummingbirds in an unchanging environment with medium flowers, the most common medium-sized bills would be favoured as longer bills may require more energy to grow and carry around and short bills limit access to food. Disruptive selection: o Selection that favours individuals with OPPOSITE EXTREMES of a trait over the individuals with the intermediate variations. o Example: If a habitat exists where flowers can be found with both small and large flowers, then hummingbirds with either long or short bills. Neither plant is ideal for the medium-sized bill, decreasing their survival and ability of reproduction Sexual selection: Differential reproductive success caused by variation in the ability to obtain mates, resulting in sexual dimorphism (males and females evolving distinct appearances and behaviour) and mating and courtship behaviours. Most common types of sexual selection are female selection and male vs. male competition. o Colouring: Male peacocks have brighter plumage than female counterparts in order to attract them as a mate o Body size: Male deer are larger than females so as to better compete during mating rituals o Physical attributes: the horns of male big-horn sheep are substantially larger than the females. Males require big horns to compete head-to-head clashes. o Mating behaviour: the male bower bird creates elaborate nesting areas or “bowers” to attract a female. The most impressive bowers are those that are largest and with the most trinkets (often blue). Disadvantages include decreased longevity as a result of the brighter colours or risky behaviour NATURAL SELECTION IN ACTION: Directional selection in Tibetan population Directional selection in a human population. Tibetan people have inhabited the Himalayan Mountains for thousands of years. Oxygen levels average at 40% of that found at sea level. Short term adaptation of people from the lowlands results in increased red blood production. This over time can be harmful as it makes the blood more viscous, causing extra stress on the heart, reduced infertility and higher child mortality. Tibetans do not exhibit high red blood cell counts, but do not have issues coping at these high altitudes. Geneticists have discovered a genetic mutation that increases the oxygencarrying capacity in their blood, which has been selected for. Antibiotic resistance in bacteria: o Antibiotic resistance means that bacteria can survive in the presence of an antibiotic o Penicillin was used to treat many infections, but today it is rarely used do to the fact that many bacteria have acquired resistance to it o This resistance has been noted in several other instances with more newly developed antibiotics. Some strains of tuberculosis are resistant to ALL known antibiotics. Treating cholera has now been reduced to a single effective antibiotic. o Resistance is thought to be a result of an existing mutation. This mutation allows the mutated bacteria to survive and then reproduce, thus creating a population resistant to the antibiotic Industrial Melanism: The Peppered moth o Evolution of this moth (the onset of melanism, darker colour) is thought to be a result of an environmental change. o Industrial melanism is the darkening of several species of moths during the recent period of industrialization in many countries o Before 1848, trees within their habitat were covered in off-white lichens, which allowed for the peppered moths to camouflage. o Due to coal mining in surrounding areas, the trees became covered in soot, and the peppered moths became easy targets for prey. The black form became common. Optional video: http://www.youtube.com/watch?v=JakdRczkmNo QUESTIONS: 1. Name the 6 steps in the process of Natural Selection? 2. What is meant by survival of the fittest? Who coined this term? 3. What is an adaptation? 4. What are the different types of selection? Provide an example for each. 5. What are three examples of selection in action? 6. What is industrial melanism? Describe the mechanism of natural selection in the peppered moth.