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Darwin’s Theory of Evolution Chapter 15 The Puzzle of Life’s Diversity Charles Darwin – THEORY OF EVOLUTION BY MEANS OF NATURAL SELECTION Made observations and devised a theory on how life changes over time Galapagos Islands and Beyond Voyage of the HMS Beagle Alfred Russel Wallace • Contemporary/ collaborator of Darwin’s • Sent a letter to Darwin summarizing similar thoughts on evolution • Darwin presented Wallace’s work to the Linnaean Society shortly before publishing On the Origin of Species by Means of Natural Selection • Credited with observing biogeographical distribution of species Darwin was influenced by several other scientists: -James Hutton and Charles Lyell – Studied rock formations and changes in the earth -Alfred Wegner - began theory of continental drift; coined term “Pangaea” -Jean Baptiste de Lamarck – Theory of Inheritance of Acquired Characteristics -Charles Lyell - considered to be the father of modern Geology. James Hutton (Scottish genius), he concluded by direct observations of rock strata that a vast amount of time was necessary for formations to form. Began idea of uniformitarianism Uniformitarianism suggested that the landscape developed over long periods of time through a variety of slow geologic and geomorphic processes Charles Lyell - Wrote Principles of Geology in which he made key advances in stratigraphy, sedimentology, and paleontology. Began idea of “faunal succession and organic extinctions”. Lyell is considered to be the “Father of modern Geology.” On this founding document of modern geology, Lyell had argued: (1) that the geologic past can best be understood in terms of natural processes we can actually observe today, such as rivers depositing layers of silt, wind and water eroding landscapes, glaciers advancing or retreating; (2) that change is slow and steady (gradualism), rather than quick and sudden; (3) that natural laws are constant and eternal, operating at about the same intensity in the past as they do today. The Present is the Key to the Past.” Alfred Wegner • Began idea of continental drift. He noticed that South America and Africa fit together like “pieces of a puzzle.” • Coined the word Pangaea or super-continent. It is believed that it came together in the Permian period and started to split apart 200 million years ago. Jean Baptist Lamarck • Lamarck is best known for his Theory of Inheritance of Acquired Characteristics • Good idea but he was incorrect! Lamarck If an organism changes during it’s lifetime in order to adapt to its environment, those changes are passed on to its offspring. He said that change is made by what the organisms want or need. Lamarck’s Theory of Evolution • Use & Disuse Organisms Could Change The Size Or Shape Of Organs By Using Them Or Not Using Them – Blacksmiths & Their Sons (muscular arms) – Giraffe’s Necks Longer (from stretching) Lamarck’s Theory of Evolution • Inheritance Of Acquired Traits – Traits Acquired During Ones Lifetime Would Be Passed To Offspring Clipped ears of dogs could be passed to offspring! Lamarck’s Mistakes • Lamarck Did NOT Know how traits were inherited (Traits are passed through genes) • Genes Are NOT Changed By Activities In Life • Populations evolve slowly… organisms do not evolve – Organisms are not given what the “need” Darwin and Lamarck They agreed on one thing: well-adapted organisms have a better chance of surviving than others Thomas Malthus British Economist. Reasoned that if the human population continued to grow unchecked, sooner or later there would be insufficient living space and food for everyone Darwin’s Observations a) Patterns of Diversity were shown b) Unique Adaptations in organisms c) Species Not Evenly Distributed i. Australia, Kangaroos, but No Rabbits ii. S. America, Llamas Darwin’s Observations • Both Living Organisms & Fossils collected • Fossils included: •Trilobites •Giant Ground Sloth of South America This species NO longer existed. What had happened to them? Vocab • Extinct species = a species is no longer living on this earth – EX: the Dodo Bird • Extant species = a species that is currently living and reproducing on earth Definition • Evolution is the slow , gradual change in a population of organisms over time Darwin’s Observations • Left unchecked, the number of organisms of each species will increase exponentially (overproduction) • In nature, populations tend to remain stable in size • Environmental resources are limited (competition) • Only the best-suited organisms survive (survival of the fittest) LIMITING FACTORS keep populations in check • • • • • • Food or water Competition for mates Space (spreads disease) Presence or absence of antibiotics Change in temperature/ climate … Remember Malthus? Darwin’s Conclusion • Production of more individuals than can be supported by the environment leads to a struggle for existence among individuals • Only a fraction of offspring survive each generation • Survival of the Fittest Darwin’s Big Ideas • Overproduction – More offspring are produced than can survive • Competition – Resources are limited; competition ensues • Survival of the Fittest – Only the individuals best suited for their environment will survive and reproduce Darwin’s Observations • Individuals of a population vary extensively in their characteristics with no two individuals being exactly alike. • Much of this variation between individuals is inheritable. Darwin’s Conclusion • Individuals who inherit characteristics most fit for their environment are likely to leave more offspring than less fit individuals • Called Natural Selection • Acts on phenotype Natural Selection 1. Individuals vary within a species 2. Individuals compete for resources in the struggle for existence 3. Fitness is the ability of an individual to survive and reproduce in its environment Natural Selection cont. 4. Fitness is the result of adaptations 5. Adaptations are inherited characteristics or traits that increase an organisms ability to survive (or increase fitness) 6. Adaptations can be physical OR behavioral 7. Individuals that are most fit are able to survive and reproduce - “survival of the fittest.” Natural Selection cont. 8. Natural Selection – results in changes in the inherited characteristics of a population. These changes increase a species’ fitness in its environment Artificial Selection • nature provides variation, humans select variations that are useful. • Example - a farmer breeds only his best livestock Artificial Selection: Roses III. Descent with Modification • Each living species has descended with changes from other species over time IV. Evidence of Evolution 1. Fossil Record 2. Biogeography (Geographic Distribution of Living Species) 3. Anatomical Homologies (Similar Body Structures) 4. Developmental Homologies (Similarities in Embryology) 5. Molecular Homologies (Similarities in DNA sequences) Evidence of Evolution A. Fossil Record Fossils are preserved remains of ancient organisms By comparing fossils from older rocks with fossils from younger layers, could see life on Earth has changed over time. Darwin view the fossil record as a record of evolution while others in his time only viewed them as preserved remains. Evidence of Evolution B. Biogeography: Geographic Distribution of Living Species – Different environments contain similar organisms with distinct differences. Ex. Flying squirrel (N. America) and sugar glider (Australia) – Alfred Wallace Evidence of Evolution C. Homologous Body structures – Structures that have different mature forms but develop from the same embryonic tissues e.g. Wing of bat, human arm, leg of turtle – Turtle Ancient lobefinned fish vestigial organs– traces of homologous organs from other species having no apparent function e.g. Appendix (digest cellulose), pelvic girdle in whales – Alligat or Bird recapitulation- structures lost during development e.g. Gill slits http://cas.bellarmine.edu/tietjen/Ec&Ev_Distance_learning/Evidence/evidence_pix.htm Mammal Homologous Structures Ontogeny recapitulates phylogeny • ontogeny is the process of embryo development (from the Greek onto, meaning "being" or "existence" and genesis meaning "origin") • phylogeny is a species' evolutionary history (from the Greek phyla, meaning "tribe" and genesis meaning "origin") Evidence of Evolution D. Developmental Homologies: Similarities in Embryology – In their early stages of development, chickens, turtles and rats look similar, providing evidence that they shared a common ancestry. E. Molecular Homologies: Similar DNA sequences • Organisms in the same taxonomic categories share similar DNA sequences and proteins. Summary of Darwin’s Theory: Evolution by Natural Selection – – – – – Variations occur within populations, and some of the variations are favorable. More offspring are produced than can possibly survive Organisms compete for resources and individuals with favorable variations are more likely to survive. Natural Selection causes species to changes over time. Species alive today are descended with modification from ancestors Survival of the Fittest!