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Measuring Evolution Two general ways to measure Evolution Microevolution: The change of populations over generations Macroevolution: Change in a species over long periods of time (Geologic time) Microevolutionary mechanisms Natural selection acts upon an individual’s phenotype. Within the lifetime of that individual, this phenotype cannot change. • Therefore natural selection operates only on populations over many generations Vocab: • Gene pool: the entire collection of alleles among a population. • Allelic frequency: The percentage of a particular allele in the gene pool of a population. – Use % b/c population size changes from gen to gen Example snapdragon population Generation Phenotype Amount Red 8 Pink 8 White 0 Red 15 Pink 6 White 3 1 5 Frequency F F’ Allele frequency The tool biologists use to see if populations have evolved. If allele frequency has not changed over generations, then population is in genetic equilibrium (not evolving) - Relate to pop growth? - Relate to succession? If allele frequency has changed over generations, then population is evolving Ways Genetic Eqilibrium can Change (Mechanisms of Evolution) 1) Genetic Drift • Change in gene pool of a small population due to chance. • Ex- Lizards: WW & Ww= non-webbed feet • ww= webbed • Pop= 10 individuals and 3 WW are killed by earthquake, what happens to allele frequency? • Allelic frequency of W will lower in this pop and w will increase. Genetic Drift 2) Gene Flow • The gain or loss of alleles from a pop by migration • Tends to reduce genetic differences b/w populations • Ex: Humans 3) Mutations • Change in an organisms DNA that creates a new allele which leads to new phenotypes. • The source of genetic variability. • Need to be recombined • Rare; take long time to develop, can reduce fitness • (e.g.) Sickle cell anemia 4) Non-random mating • Selection of mates other than by chance. Selective breeding by populations (most do this). • Ex. Herd of elk, elephant seals, peafowl, red cross-bills. 5) Natural Selection • The factor that is likely to result in adaptive changes in the gene pool. • Acts upon the ranges of variation Pattern of Inheritance? • 3 types 1) Stabilizing Selection: • Favors the average individual in a population for a particular environment. • Spider size; small= Can’t catch enough prey • Large= easy to see and eaten • Medium= Favored • Variation is reduced • Human babies born at an average mass are more likely to survive than babies born either much smaller or much larger than average. Copyright Pearson Prentice Hall 2) Directional Selection • When one of the extreme forms of a trait is favored. • Ex. Wood pecker beak size: Short, Med, Long • In this case, birds with larger beaks have higher fitness. Therefore, the average beak size increases. Copyright Pearson Prentice Hall 3) Disruptive Selection • When both extreme forms of a trait are at an advantage • Ex: Limpets shell color white & Dark Brown • On light colored rocks white shell adv. • On darker rocks dark brown adv. • But tan shells easy to see regardless of rock color, most preyed upon. • ** can lead to the evolution of 2 new species • If average-sized seeds become scarce, a bird population will split into two groups: one that eats small seeds and one that eats large seeds. Copyright Pearson Prentice Hall