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Standard 5 Unit 12 – Evolution & Biodiversity Objective B5.1A: Summarize the major concepts of natural selection (differential survival and reproduction of chance inherited variants, depending on environmental conditions). B5.1c: Summarize the relationships between present-day organisms and those that inhabited the Earth in the past (e.g., use fossil record, embryonic stages, homologous structures, chemical basis). Theory of Evolution A. Idea supported by scientific evidence (but no concrete experiments) over a long period of time B. Change in a species due to mutation of the DNA code that occurs over a long time. Evolution is the result of genetic changes that occur in constantly changing environments. Evolution is all the changes that have transformed life from its earliest forms to what we see today. Working definition = Evolution is the progressive change in organisms over time. C. D. E. Evolution of Air Breathing: Evolution of Brains: Charles Darwin Natural Selection: “Survival of fit” Fit reproduce Competition for resources Best adapted species survive Introduction to Charles Darwin's Theory of Evolution (00:54) HMS Beagle Voyage 1835 Darwin’s Voyage of Discovery A reconstruction of the HMS Beagle sailing off Patagonia. The 5 Year Voyage of the Beagle Darwin’s key ideas: A. REPRODUCTION: Organisms produce more offspring than can survive B. VARIATION:Variety in traits exist C. SURVIVAL OF THE FIT: Some traits allow survival & are passed on D. Over time certain variations make up most of a population & they may be different from their ancestors Darwin’s Ideas Did Not Develop in a Vacuum Contributor’s to Darwin’s thinking included: – Charles Lyell – uniformatarianism – Georges Cuvier – species extinction – Thomas Malthus – struggle for existence – Jean Baptiste de Lamarck – evolution by acquired characteristics. – Alfred Russel Wallace Independently Drew the Same Conclusions as Darwin Malthus’s contribution: Populations grow to a maximum level Environmental limitations Fit animals out compete the less fit P O P U L A T I O N 50% 30% Carrying Capacity 10% 10 yr 20 yr 40 yr 60 yr 80 -10% Darwin’s Observations & Inferences Observation 1: Left unchecked, the number of organisms of each species will increase exponentially, generation to generation. Observation 2: In nature, populations tend to remain stable in size Observation 3: Environmental resources are limited. Observation 4: Individuals of a population vary extensively in their characteristics with no two individuals being exactly alike. Observation 5: Much of this variation between individuals is heritable. Darwin’s Observations & Inferences Inference 1: Production of more individuals than can be supported by the environment leads to a struggle for existence among individuals, with only a fraction of offspring surviving in each generation. Inference 2: Survival in the struggle for existence is not random, but depends in part on the heritable characteristics of individuals. Individuals who inherit characteristics most fit for their environment are likely to leave more offspring than less fit individuals. Inference 3: The unequal ability of individuals to survive and reproduce leads to a gradual change in a population, with favorable characteristics accumulating over generations (natural selection). Taken together, these three inferences are a statement of Darwin’s Theory of Evolution. Evolution Evidence: 1. Adaptations 2. Fossils 3. Comparative anatomy 4. Comparative embryology 5. Comparative Biochemistry 6. Plate Techtonics 1. Adaptations: features suited to a particular environment that allow organisms to survive Inuit people, who live in the extreme cold of the Arctic, have short, stout bodies that conserve heat. Adaptations Masai people, who live in the arid lands of eastern Africa, have tall, lean bodies that disperse heat well. Plant Adaptations: Help!!! Venus Fly Trap Captures Animals Acquires Minerals For Photosynthesis Leaf Adaptations: Succulents Thick Store Water Prevent Drying out Leaf Adapatations: Pine Needles Shed snow Less water loss Reduced surface area Tolerate wind Flower Adaptations: Fly pollination: Hair along petals Putrid smell Bee pollination: Smooth petal Sweet smell 2. Fossil Evidence: The Evidence for the Theory of Evolution (02:04) 1. 2. 3. Once living remains of organisms Limited to: Type of material preserved (bone, shell, impressions, amber) Incomplete record Easily disrupted Plant Fossil Evidence: 3. Comparative Anatomy: Structural similarities link related species Comparative Anatomy Structures: Analogous: 1. Different ancestors 2. “analogy”=like 3. Different underlying structures 4. Same Function 5. Similar Environments Homologous: 1. Same ancestor 2. “homo”=same 3. Same underlying structures 4. Different Functions 5. Different Environments Analogous Structures Different underlying structures (different ancestors) Same function, similar environments Homologous Structures: Same underlying structures, different functions, different environments & common ancestor 4. Comparative embryology: Similar embryo development in closely related species 5.Comparative Biochemistry Similar DNA sequences= Similar Gene segments of the DNA Code for similar traits In closely related species Kinship There is a degree of kinship between organisms or species that can be estimated from the similarity of their DNA and protein sequences. Chimpanzees are so closely related to humans that they should properly be considered as members of the human family, according to new genetic research. Scientists from the Wayne State University, School of Medicine, Detroit, US, examined key genes in humans and several ape species and found our "life code" to be 99.4% the same as chimps. They propose moving common chimps and another very closely related ape, bonobos, into the genus, Homo, the taxonomic grouping researchers use to classify people in the animal kingdom. Humans, or Homo sapiens to give the species its scientific name, are the only living organism in the genus at the moment although some extinct creatures such as Neanderthals (Homo Neanderthalis) also occupy the same grouping. Dr Wildman said: "You could say that humans and chimps are as similar to one another as say horses and donkeys. Evidence of Evolution – Conservation and Diversification at the Molecular Level Why should different organism possess related genes? Why does the degree of relationship of genes match their degree of relationship established by other methods? Evidence for Evolution – Evolution Observed Evolution of pesticide resistance in response to selection. Evidence for Evolution –Evolution Observed Evolution of drug-resistance in HIV 6. Plate Techtonics Geological theory: Continental masses were one land mass that explains Closely related species have common ancestors on now separated continents What Darwin Never Knew http://video.pbs.org/video/137207 3556/ Movie Trace the relationship between environmental changes in the gene pool, such as genetic drift and isolation of subpopulations. Biomes: Islands and Evolution (56:00) Early Theories of Evolution: Darwin: Current theory Natural Selection “Survival of fit” Reproduction of the best adapted species Lamark: “Use & Disuse” Abandoned No knowledge of genetic traits or mutations in sex cells Lamark’s Theory “Use and Disuse” Use of structure results in evolution Does not take into account DNA or sex cell mutations I. What is evolution? A change in the genetic makeup of a population with time. It is a two-step process – Development of genetic variability in the population. – Natural selection of those variations which are favorable for survival. Example Let’s say a population of black spotted cattle--BB-- eventually turn into a population of white spotted cattle--bb-- over a period of years. II. Genetic variability Assuming that a population starts out with the same genetic makeup, all new genetic material in that population will arise by mutation. – A mutation is a random inheritable event. – Let’s take our black spotted cattle as an example; for some reason the chromosomes that code for black spots are mutated. Variability This mutation Changes the BB to Bb. Now with the b gene in the herd, White spotted cattle can arise. Variation Variation in the chromosomes occurs when an egg and a sperm unite. Both carry unique traits from the parents into one individual, forming an individual which varies from the parents. – This variation is normally a random event. – In our example, let’s say two mutated animals mate with one another. Let’s look at their offspring. B B BB (black) b Bb (black) b Bb (black) bb (white) A Scenario to demonstrate natural selection It has been 50 years since the first mating of our mutated cattle. Over the years the once all Black spotted (BB) herd is now about 75% black spotted and about 25% white spotted (bb). A new disease has mutated during this time period. A disease known as Bovine Black Spotitis. The disease is not fatal, but any black spotted (BB or Bb) bull or cow is sterile. Question? The disease spreads quickly through the herd and within six months time all black spotted cattle are sterile. What will eventually happen to the herd? Answer Only the white cattle will be able to reproduce, eventually producing a completely herd of white spotted cattle. (bb) Natural Selection As long as the environment does not again change then these individuals will become the “normal” population. The sequence of events that leads to a certain characteristic being “selected” in the environment is called natural selection. Populations change in response to their environment. Acts on phenotype of an organism. (Not genotype) Determines the differential survival of groups of organisms Evolution’s Core Principles Natural selection Evolution’s Core Principles Common descent with modification Macroevolution: Long time scale events that create and eliminate species Variations: Differences in traits Come about by mutations in genes Random Occur in sex cells Passed on to future generations Bird Beak Adaptations: Genetic Drift Changes in the gene pool due to: 1. 2. 3. 4. 5. Random mating Over a long time period No immigration of males No emigration of females Sufficient resources that match the adaptations Same Species Must: Show similar characteristics Successfully interbreed Producing fertile offspring Donkey + Horse= Mule (infertile) Speciation Evolution New Species Over time By Isolation Natural Barriers Speciation Formation of a new species Geographic Isolation Separation of organisms by geographic features. A physical barrier divides a population: – Mountains – Lakes, oceans, rivers – Desserts These two groups are not able to mate => may lead to new species Reproductive Isolation When two different species can not mate and have successful offspring – Geographic barriers – Anatomy or physiology – Social behaviors Prevention of mating between formerly interbreeding groups. or The inability of these groups to produce fertile offspring Reproductive Isolation: Two organisms cannot mate Separated by geographic boundaries Anatomical differences Physiological differences Social behaviors Gradualism “gradual” Small changes Over a long time Gradualism Model of evolution involving slow, gradual changes to a species over time. Punctuated Equilibrium “punctuation!” Large changes Happen rapidly Periods of no change Punctuated Equilibrium Evolutionary model Rapid changes to a species in a short time, followed by periods of little change. Gradualism vs. Punctuated Equilibrium Gradualism: # S P E C I E S Time Punctuated Equilibrium Gradualism vs. Punctuated Equilibrium Adaptive Radiation: “radiation”= branching from one source “adaptive”= survival of fit Evolution of many branches of organisms from a single source Adaptive Radiation Divergence “diverge”= branch off Homologous structures Same origin Same underlying structure Difference functions Human arm Bat wing Cat limb Whale flipper Original Species: Mammal Convergence Bird wing “converge”=come together” Analogous features From different origins Organisms that fly Similar environments= Similar functions Different structures * Homework – 43-1 Butterfly wing Bat wing Convergent Evolution Placental mammals Marsupial mammals Convergent Evolution Although marsupial mammals once populated all land masses, they remain diversified only on the isolated Australian continent, where they have evolved to fill the same ecological niches that placental mammals occupy elsewhere. The Tasmanian wolf, for example, closely resembles the doglike carnivores of other continents. More specialized parallel adaptations include those of the marsupial and placental anteaters, the marsupial sugar glider and placental flying squirrels, and the burrowing marsupial wombat and placental ground hog. In this illustration, placental mammals are in the top row, and their marsupial equivalents are in the bottom row. Genetic Variation * Homework – Give examples of ways in which genetic variation and environmental factors are causes of evolution and the diversity of organisms.