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Evolution of Populations Chapter 11 Warm Up 1/21 & 1/22 1. Compare Larmark and Darwin in their theories of evolution. 2. List the 5 evidences of evolution. 3. Define the following: Vestigal, fitness, mutation and adaptation KEY CONCEPT A population shares a common gene pool Genes and Variation Gene pool - all the genes that exist within a population Genetic variation in a population increases the chance that some individuals will survive Genetic variation leads to phenotypic variation. Phenotypic variation is necessary for natural selection. Genetic variation is stored in a population’s gene pool. made up of all alleles in a population allele combinations form when organisms have offspring Allele frequencies measure genetic variation. measures how common allele is in population can be calculated for each allele in gene pool Genes and Variation Relative (allelic) frequency - the percentage of a particular allele in a gene pool. KEY CONCEPT Populations, not individuals, evolve. Natural Selection in Populations Natural Selection- In nature, unequal ability to survive and reproduce... Survival of the fittest. Natural Selection ACTS ON PHENOTYPE but influences genotype (thus, allelic frequency) Artificial Selection- Mankind selects for desired traits. Also known as “selective breeding” Over time, the zebra herd becomes faster as the slower zebra (and their genes) are removed from the population… survival of the fittest Darwin’s Theory = Evolution by means of natural selection ADAPT OR DIE! Camouflage- organisms blend-in with surrounding environment Mimicry- species copy another to insure their own survival NatGeo Natural Selection effects Genetic Change in Populations Natural Selection has three affects on phenotype distribution 1.Directional Selection 2.Stabilizing Selection 3.Disruptive Selection Artificial Selection Normal Distribution of traits A normal distribution graphs as a bellshaped curve. Phenotypes near the middle range tend to be most common. Examples- height and weight Directional Selection This type of selection favors phenotypes at one extreme of a trait’s range. An extreme phenotype that was once rare becomes more common. Ex. Drug resistant bacteria Directional Selection 1. Directional SelectionIndividuals on one end of a curve are “better fitted” than the middle or other end Peccaries naturally choose to consume those cactus plants with the fewest spines As a result, at flowering time there are more cacti with higher spine numbers; thus, there are more of their alleles going into pollen, eggs, and seeds for the next generation. Stabilizing Selection Stabilizing selection favors the intermediate phenotype. Selection against both extremes decreases the genetic diversity of a given population. Stabilizing Selection 2. Stabilizing SelectionIndividuals near center of a curve are “better fitted” (have highest fitness) than both ends Peccaries are consuming the low-spine number plants, and the insects are killing the high-spine-number plants. As these gene combinations are removed from the cactus gene pool, there is less and less variety possible in subsequent generations. Disruptive Selection This type of selection occurs when both extreme phenotypes are observed. Individuals with the intermediate type are selected against. By favoring both extreme phenotypes, disruptive selection can lead to the formation of a new species. Disruptive Selection 3. Disruptive SelectionIndividuals at upper and lower ends are “better fitted” than the ones in the middle Years of collecting have left their toll on the roadside cacti. In this environment, it is maladaptive to be good looking and have a reasonable number of spines. Low spine-number plants are not picked because they don't "look right", and high spine-number varieties are left alone because they are too hard to pick. Gradually, the gene pool changes in favor of the two extreme spine number types. Other mechanisms of Evolution Natural selection is not the only mechanism through which populations evolve 3 other mechanisms of Evolution: Gene Flow Genetic Drift Sexual Selection Gene flow – movement of alleles into or out of a population Immigration – new alleles move IN Emigration – alleles move OUT Genetic drift - change in allelic frequencies by chance Ex: sudden extinction of a dominant species; small populations most affected Genetic drift is a change in allele frequencies due to . chance Genetic drift causes a loss of genetic diversity It is most common in small populations. A population bottleneck can lead to genetic drift. It occurs when an event drastically reduces population size. The bottleneck effect is genetic drift that occurs after a bottleneck event. The founding of a small population can lead to genetic drift. – – It occurs when a few individuals start a new population. The founder effect is genetic drift that occurs after start of new population Sexual selection occurs when certain traits increase mating success. Sexual selection occurs due to higher cost of reproduction for females. males produce many sperm continuously females are more limited in potential offspring each cycle Genetic equilibrium - when alleles stay the same from generation to generation 1. 2. 3. 4. 5. The Hardy Weinberg Principle: Allele frequencies will remain constant under five conditions Random Mating Large Population No movement (immigration or emigration) No Mutations No Natural Selection: equal change of survival 5 Factors that can lead to evolution Genetic Drift Gene Flow Mutation Sexual Selection Natural Selection Genetic drift changes allele frequencies due to chance alone Gene flow moves alleles from one population to another Mutations produce the genetic variation needed for evolution. Sexual selection selects for traits that improve mating success. Natural selection selects for traits advantageous for survival KEY CONCEPT New species can arise when populations are isolated The isolation of populations can lead to speciation speciation - evolution of a new species . Reproductive isolation can occur between isolated populations Populations can become isolated in several ways: 1. Behavioral 2. Geographic 3. Temporal 1. Behavioral Isolation: Two populations capable of breeding but cannot because of courtship rituals 2. Geographic Isolation: Two populations are separated by geographic barriers Ex: Rivers, Oceans, Mountains 3. Temporal Isolation: Two or more populations reproduce at different times Patterns of Evolution 1. 2. 3. 4. Extinction Divergent Evolution (adaptive radiation) Convergent Evolution Coevolution 1.Extinction Why do species go extinct? Extinction Natural selection, climate changes, and catastrophic events have caused 99 percent of all species that have ever lived to become extinct. Mass extinctions – caused by continents moving, sea level changing, volcano eruptions, large meteors Predict what each ecosystem will look like after the event. Catastrophic Event Catastrophic Event Question When a mass extinction happens, what do you think will happen next? Divergent Evolution (adaptive radiation) Divergent evolution – natural selection causes 1 species to evolve into many species with many different adaptations (homologous structures) After mass extinctions, many environments will be open for inhabitation Species will migrate to that area and new environmental pressures will cause the population to change over time This is also known as Adaptive Radiation Adaptive Radiation in honeycreepers Evidence of Evolution Homologous structures - similar structures found in related organisms that are adapted for different purposes. Ex: human arm and bat wing or whale flipper ---DIVERGENT EVOLUTION--the process of two or more related species becoming more and more dissimilar. Homologous structures Divergent evolution Convergent Evolution Convergent Evolution – when unrelated organisms come to resemble one another (analagous structures) Analogous structures - structures found in unrelated organisms that have a similar function but may be structurally different Ex: bird wing and insect wing ---CONVERGENT EVOLUTION--independent evolution of similar features in species of different lineages Analogous structures Convergent evolution Coevolution When 2 species evolve in response to one another Coevolution can occur in competitive relationships, sometimes called evolutionary. Speciation occurs in patterns Punctuated equilibrium: species show little evolutionary change for millions of years, followed by periods of rapid speciation Gradualism- Species evolve slowly, over long periods of time.