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Genetic variation and Evolution Chapter 20 Why are organisms well adapted to their habitats? Creationist explanation (see section 21.7) 20.2 -- should know H-W equation and why it’s important -- we may come back to this Is a religious view Divine role 20.4 – understand the meaning of fitness, but we’ll not cover the mathematical aspects 20.5 – skip “intelligent design” not scientifically testable 20.6 – read on your own 20.8 – read on your own Scientific explanation “evolution from common ancestor” ‘Natural Selection” -- is testable Question Read or listen to the The Firing Line 1997 Creation-Evolution Debate http://www.bringyou.to/apologetics/p45.htm Genetic Variation & Evolution 1 Darwin was not first to describe evolution George Cuvier (1790) – Fossils & ‘Catastrophism’ Jean Lamarck (1809) -- Environmental influence Lamarck’s theory of “Inheritance of Acquired Traits” Question Genetic Variation & Evolution 2 Modern explanations of why traits change over time (20.3) -- Frequency of genes in the population change 5 Agents (mechanisms) of change: 1. Genetic Mutation – random changes to DNA code -- creates variation in traits -- ‘raw material’ of all evolutionary mechanisms 2. Natural selection – mechanism proposed by Darwin -- environment selects certain traits Genetic Variation & Evolution 3 Agents of change, con’t Gene Flow Other agents of change 3. Gene flow -- movement of individuals between populations 4. Nonrandom mating -- organisms prefer to mate with certain individuals 5. Genetic drift -- chance may favor certain traits in a small population Founder effect Bottleneck effect Founder effect – population isolation bottleneck effect – population catastrophe -- gene pool reduction Genetic Variation & Evolution 4 What are the fundamental premises of Darwin’s theory of “Natural Selection”? Read Section about Evolution in Chapter 1 “On the Origin of Species by Means of Natural Selection” “Evolution” (a process) does not equal “Natural Selection” (a process) Natural selection occurs when: 1) There exists genetically based variation (inheritable) in the traits of a species. 2) Organisms produce more offspring than can survive. 3) The environment selects for those organisms with the best adapted traits. 4) Organisms best adapted to the environment will reproduce most successfully. Genetic Variation & Evolution 5 How does Darwin’s ‘Natural Selection’ explain giraffe evolution? “Selective Pressures” “Adaptation” -- a noun and verb -- not an ecological term Genetic Variation & Evolution 6 What is nature of genetic variation? (20.1) Genetic mutation DNA Polymorphisms Genetic variation in Asiatic lady beetle Different ‘alleles’ Hardy-Weinberg equation (20.2) p2 + 2pq + q2 -- frequency of Alleles frequency of traits -- can provide evidence for selection Question Genetic Variation & Evolution 7 What are examples of environmental selective pressures? Predator-prey interactions Competition for resources Human impacts Natural selection acts upon existing genetic variation -- theory is not ‘teleological” Wrong: “In order to hide from predators, the dead-leaf butterfly evolved a leaf-like appearance.” Better: “Predation selected for butterflies that most resembled a dead leaf.” The environment selects among but does not create variations! Question Genetic Variation & Evolution 8 Do organisms always produce more offspring than can survive? Cichlides: 250,000 eggs Carmagnola Grey rabbit: 5-8 litters x 7 offspring Genetic Variation & Evolution Humans: 2- 7 children/family 9 Why don’t humans evolve wings? Limits of Selection Existence genetic variation Complex traits involve many genes Suitable selective pressures Genes have multiple effects -- developmental genes Question Genetic Variation & Evolution 10 What is ‘Artificial Selection’? All these crops are varieties of the species Brassica napa Are humans still evolving? Colors represent different genetic types. Map shows genetic variation in Africa. Group project Genetic Variation & Evolution 11 Why does ‘in breeding’ cause genetic problems? species non-inbred inbred Zebra lived died 20 7 3 2 Eld’s deer lived died 13 4 0 7 Giraffe lived died 11 3 2 3 Oryx lived died 35 2 0 5 Dorcas gazelle lived died 36 14 17 25 From Klug & Cumminings Genetics 5th ed Genetic Variation & Evolution 12 What are the effects of Natural Selection on a population? (20.7) “Microevolution” vs “Macorevolution” Genetic Variation & Evolution 13 What is the effect of Directional Selection on a population? Progressive change in a trait within a species Genetic Variation & Evolution 14 How does Stabilizing Selection affect a population? Human birth weight & Eurosta gall size Stabilizing selection & ‘living fossils’ Genetic Variation & Evolution 15 How does Disruptive Selection affect a population? Responsible for divergence of a single species into two different species Modern example of process: blackbellied seedcracker Question Genetic Variation & Evolution 16