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Angiosperms III Plant Cell Types Plant Cell Types • Most angiosperm plants are composed of 10 fundamental cell types • Many more specialized cell types do occur in some plants • These 10 cell types are differentiate from the dividing cells found in meristems and cambia (meristematic cells) Meristematic cells • Found in meristems (root and shoot), the cambial layers, and buds • Function is cell division (cell cycle) • Small (about 10 m) with a large nucleus (the cells you saw in the squashed root tip in the mitosis lab) Parenchyma Cells • • • • Large (100m) cells Thin cellulose cell walls Usually with a large vacuole Function as storage or “packing” cells • Common in the pith and cortical layers, the wood rays, petals, etc… • A “generalist” cell type Chlorenchyma Cells • Large parenchyma cells with chloroplasts • Found in the leaves and in some stems • Major function is photosynthesis Collenchyma Cells • Cells used for temporary support, primarily in herbaceous stems • Cells walls have extra cellulose thickenings in the corners (strong but flexible) • Often found in bundles as “collenchyma tissue” just under the epidermis • Best experienced in celery (or rhubarb) as the “ribs” that get stuck in your teeth Sclerenchyma Cell Types • “Sclerenchyma cells” are four cell types which all share these features in common: – all have thick secondary cell walls of LIGNIN (and stain red in prepared slides) – all are DEAD at maturity – include: fibers, sclereids, tracheids, and vessels elements Fibers • Long, narrow cells with thick walls and a small lumen • Used for support in woody tissues (secondary xylem) or for support in vascular bundles and herbaceous stems • Not found in conifer wood (thus the wood is “soft”) Fibers (longitudinal section) Sclereids • Irregular cells with thick walls • Function primarily for protection (surfaces often look like jigsaw puzzles) • Found in dry fruits (nuts etc.), seed coats, some leaves Sclereids • A special form of a sclereid, the stone cell, gives pears their “gritty” texture Tracheids • Water conducting cells of the xylem in gymnosperms, ferns, and fern allies • Cells have closed end walls • Found along with vessels in angiosperms • May have complex “pits” through which water moves from cell to cell • Secondary wall may be deposited in rings or spiral shapes Vessels Elements • Water conducting cells of the xylem in angiosperms; also found in gnetophytes • Have open end walls • May have spiral or ringlike secondary cell wall patterns Vessel Elements • Are short and squat cells, one on top of another, forming vessel “tubes” for conduction Sieve Tube Members • Tall, thin cells with only primary cellulose cell walls • Living cells but lack a nucleus at maturity • Similar to sieve cells of gymnosperms Sieve Tube Members • Part of the phloem tissue which conducts sugars and hormones throughout the plant Sieve Tube Members • End walls have a “sieve plate” Companion Cells • Long, thin cells with a nucleus directly adjacent to each sieve tube members • Sieve tube member and its companion cell are derived from the same initial • Responsible for the loading and unloading of substances from the sieve tube members in the phloem • Similar to albuminous cells in gymnosperms Development of Sieve Tubes and Companion Cells Summary of Cell Types • • • • Meristematic Parenchyma Chlorenchyma Collenchyma • Fibers • • • • Sclereids Tracheids Vessel Elements Sieve Tube Members • Companion Cells