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Unit #1: Evolution Natural selection can change gene frequencies through the differential survival and reproduction of organisms that are better adapted to their environment (more fit) (LO 1.5, LO 1.2, EK 1.A.A, EK 1.A2 Natural selection can be disruptive, directional, or stabilizing (LO 1.22 Adaptations are heritable traits that increase reproductive success and survival Mutations provide variation in traits that selecting agents act on Natural selection is the only evolutionary force that is adaptive Sexual selection is a type of natural selection Natural selection does not make perfect organisms At the most basic level, evolution is a change in gene frequencies (the proportion of each allele in a gene pool) in a population from one generation to the next ∫EU 1.A Artificial selection can change gene frequencies when humans manipulate selecting agents in the environment Antibiotic and pesticide resistance are two examples Lab 1: Artificial Selection Mutations are random changes in DNA that can change gene frequencies Gene flow can change allele frequencies through the migration of new individuals into or out of a population Genetic drift can change gene frequencies through chance events EK 1.A.3, LO 1.8 Mutations occur spontaneously, due to errors in DNA replication or mutagens EK 1.A.3 Founder’s effect occurs when part of population colonizes a new area Bottleneck occurs when the size of a population is drastically reduced by disaster Mutations do not occur in response to a need in environment Some mutations are positive, some are negative, and some are neutral Whether a mutation is + or - depends on the environment LO 1.5 Unit #1: Evolution The Hardy-Weinberg equation provides a mathematical model for analyzing evolutionary change by measuring and predicting changes in gene frequencies ∫EU 1.A P2, 2PQ and Q2 represent the frequencies of homozygous dominant, heterozygous and homozygous recessive genotypes in a gene pool, while P and Q represent the frequencies of the dominant and recessive alleles in a gene pool. (LO 1.3, LO 1.6, LO 1.7 The sum of the frequencies of all genotypes in a gene pool must equal 1, while the sum of the frequencies of the alleles in a gene pool must equal 1. (LO 1.3, LO 1.6, LO 1.7 Genomes reveal both neutral and selective processes of evolution. Heterozygotes maintain diversity in pool HW equilibrium occurs when gene frequencies do not change from one generation to the next. Conditions for equilibrium include no gene flow, no drift, no mutation, no selection, and random mating (LO 1.1) Conditions for equilibrium are rarely met Lab 2: Mathematical Modeling Quiz: Microevolution Evolution is still occurring in populations – finches, chemical resistance, diseases, sickles cell, eukaryotic st. (EK 1.C.3, LO 1.25, LO 1.26) Heterozygote advantage occurs when heterozygous genotypes are more adaptive than others Recombination, lateral gene transfer and gene duplication can result in new features A diverse gene pool can aid in the survival of a population Much of molecular evolution is neutral Recessive mal-adaptive alleles are rarely eliminated from gene pool because of heterozygotes. Genome size and organization can evolve Unit #1: Evolution All life descends from one common ancestor ∫EU 1.A There are a variety of sources of evidence for evolution: anatomical, fossil, biogeographical, molecular (EU 1.B) Comparison of DNA and amino acid sequences provides molecular evidence for divergent evolution from a common ancestor All cells have DNA, RNA, metabolic processes, gene sequences (LO 1.14, LO 1.15, LO 1.16) Much of molecular evolution is neutral Genome size and organization can evolve Comparison of anatomical features such as homologous structures, vestigial structures, and embryological development provide evidence of divergent evolution from a common ancestor Analogous structures are evidence of convergent evolution due to similarities in the environment but not a recent common ancestor Fossils provide evidence of species that no longer exist and provide a means of relatively dating evolutionary events Fossils can be dated relatively by stratigraphy or quantitatively by radiometric dating and paleomagnetic dating Phylogenetic trees show divergence of species from a common ancestor the evolutionary history of species Molecular and anatomical data can be used to build and test phylogenetic trees (LO 1.11, LO 1.12, LO 1.13) Phylogenetic trees can show homologies that are derived or lost due to evolution Molecular clocks date evolutionary events Lab 3: DNA Sequences & Evolutionary Relationships using BLAST Unit #1: Evolution New species evolved and continue to evolve within a changing environment EU 1C, EK 1C3 Speciation occurs when two populations become so different genetically that they can no longer interbreed successfully EU 1C, EK 1C3 ∫EU 1.A Allopatric speciation involves both geographic isolation and reproductive isolation (LO 1.23) EK 1C2 Sympatric speciation can occur without geographic isolation through polyploidy and sexual selection ∫EU 1.A Because of life’s diversity, determining which groups of organisms represent distinct species can be complex Morphological species concept is limited; Biological species concept is used more widely Pre and post zygotic mating barriers enforce reproductive isolation and the accumulation of genetic differences Speciation can occur at different rates (punctual equilibrium and gradual) LO 1.23 Adaptive radiation is divergence of many new species from one ancestral species due to different environments LO 1.20, LO 1.21 Major changes in earth’s physical environment such as cont. drift, climate, volcanoes, meteors, and oxygen have affected the evolution of life When the environment changes, there is no guarantee that a species will adapt LO 1.23 Speciation is still occurring Gene flow can change frequencies through migration of new individuals There have been several mass extinctions in history LO 1.20, LO1.21 Extinction eliminates species but can also provide opportunities for new species to arise Hybrid zones can form if reproductive isolation is incomplete Quiz: Macroevolution Extinctions are still occurring today, often due to human impact Unit #1: Evolution Evolution can explain the origin of the first life forms EU 1D, EUK 1.D.1, EK 1.D.2∫EU 1.A Evidence shows that earth is 4. BYA and the first life evidence is 3.5 BYA All life is related through a common ancestor according to genome sequences The molecular building blocks of life were present on earth before life evolved but not O2 LO 1.27, 1.28, 1.29, 1.30, 1.31, 1.32 Spontaneous chemicals reactions in the absence of oxygen resulted in the formation of increasingly complex organic molecules from inorganic molecules (organic soup) Scientific experiments have simulated conditions that could explain the evolution of first life forms (organic from inorganic) LO 1.27, 1.28, 1.29, 1.30, 1.31, 1.32 LO 1.23 Lab 4: Organic Soup Model RNA may have been the first genetic material Unit Exam All cells have DNA, RNA, metabolic processes, gene sequences (LO 1.14, LO 1.15, LO 1.16) Eukaryotes have structural similarities cytoskeleton, organelles, linear chromosomes Events can be dated relatively by stratigraphy or quantitatively by radiometric dating and paleomagnetic dating Evolution Unit Plan Description: In this unit, students will explore mechanisms of both microevolution and macroevolution. Students will first investigate natural selection as a force of evolutionary change that results in populations better adapted to their environment, and then contrast selection with other evolutionary forces such as gene flow, drift and mutation. Students will measure, analyze and predict evolutionary change using mathematical models. Students will also examine a range of evidence that supports evolutionary theory, and use that evidence to create, evaluate and justify phylogenetic trees. Students will then analyze large-scale evolutionary processes, including speciation and extinction. Finally, students will describe and evaluate hypotheses for the origin of the first life forms. The unit will draw on and connect prior knowledge of genetics, biochemistry, cellular biology, anatomy and reproduction. Enduring Understanding: Evolution drives the unity and diversity of life / Evolutionary theory explains the descent of millions of species from one common ancestor. Essential Questions: How do we explain the current diversity of life on earth? Essential Sub-Questions How and why do populations evolve? How do groups of organisms become adapted to their environment? Is evolution progress? How do we know evolution occurred in the past, and how do we know it is occurring today? How do new species arise? Did humans come from monkeys? Could humans evolve into a new species? Where did the first cell come from? Learning Goal Evolution is a change in the genetic makeup of a population over time as a result of several possible forces. Sub-learning goals Evidence An adaptation is a genetic variation that results in a trait that provides an advantage to an organism’ survival in a particular environment. Phenotypic variations are not directed by the environment but occur through random changes in DNA and through new gene combinations. Evolutionary fitness is measured by reproductive success. Natural selection is an evolutionary force that produces betteradapted populations through competition for limited resources that results in differential survival and reproduction. The environment affects evolutionary rate and direction by acting as a selecting agent for adaptations. There are different types of natural selection, including directional, stabilizing, disruptive and sexual. Humans act as selecting agents and impact variation in other species through artificial selection. Chance and random events such as genetic drift, gene flow and mutation act as nonselective evolutionary forces. Genetic drift includes the bottleneck effect and founder’s effect. Mutations can be positive, negative or neutral – often depending on the environment. Mutations, gene transfer and recombination through sexual reproduction help maintain a diverse gene pool, which in turn can increase the ability of a population to survive. Conditions necessary for a population to remain at equilibrium include large population size, absence of migration, no net mutations, random mating and no selection. Conditions for equilibrium are rarely met. Mathematical models such as the Hardy-Weinberg equation can be used to calculate changes in gene frequency and provide evidence for evolution. Natural selection rarely eliminates harmful recessive alleles because the heterozygote genotype masks the allele. Heterozygote advantage occurs when individuals with the heterozygote genotype are more fit than homozygous dominant or recessive individuals. Misconceptions Explain how natural selection leads to better adapted populations Compare the effect of various types of natural selection (including stabilizing, directional, diversifying and sexual) on the genetic makeup of a population Design, conduct, and analyze results of an artificial selection experiment (Lab 1) Describe and differentiate the effect of nonselective evolutionary forces on a population Identify conditions necessary for Hardy Weinberg equilibrium Use the Hardy-Weinberg equation to calculate gene and genotype frequencies, determine whether a population is evolving or in equilibrium, and predict changes in the population Explain how heterozygote advantages occurs in sickle cell anemia Manipulate and analyze data for an evolving population using a computer spreadsheet and then justify conclusions using data (Lab 2) Individual organisms can evolve Mutations/adaptations occur in response to environmental changes Mutations are always harmful Evolution always produces betteradapted organisms Natural selection will eliminate harmful alleles from a population Natural selection makes perfect organisms A variety of evidence supports the claim that all life descends from a common ancestor All organisms share certain conserved core process and features, including DNA, RNA, and metabolic pathways All eukaryotic organisms share certain structures, including cytoskeleton, membrane-bound organelles, linear chromosomes and endomembrane systems Morphological homologies, including vestigial structures, provide evidence of descent from a common ancestor (divergent evolution) Biochemical and genetic similarities, specifically amino acid and nucleotide sequence, provide evidence of descent from a common ancestor (divergent evolution) Fossils can be dated and provide evidence of species that no longer exist. Analogous structures have similar functions but different composition and provide evidence of convergent evolution of species not closely related but found in similar environments. Phylogenetic trees are graphical representations or models of evolutionary history Phylogenetic trees can be constructed based on morphological and molecular evidence Phylogenetic trees can be tested and revised Phylogenetic trees can show traits that are derived or lost due to evolution Phylogenetic trees show speciation events, common ancestry and relatedness of different groups Computer programs have sophisticated methods of measuring molecular data and representing relatedness among organisms Describe examples of conserved core biological processes and features share by all domains of life and within one domain of life Differentiate and relate morphological, molecular, fossil and biogeographical evidence for evolution Justify the claim that life descends from a common ancestor using multiple forms of evidence Infer evolutionary relationships of groups shown in phylogenetic trees and evaluate the evidence for those relationships Construct and evaluate phylogenetic trees using morphological data Use BLAST to compare genes from different organisms and construct a cladogram to model the evolutionary relatedness of the species (Lab 3) Evolution is “just” a theory Evolutionary trees are static Humans evolved from monkeys New species have formed and continue to form within a changing environment. Speciation is the formation of a new species and results in diversity of life forms Speciation occurs when two groups of organisms become so different genetically that they can no longer interbreed successfully Speciation can occur when geographic isolation is followed by reproductive isolation, which may take hundreds, thousands or millions of years. Speciation can occur rapidly and without geographic isolation through polyploidy Various pre- and post-zygotic matting barriers can maintain reproductive isolation and prevent gene flow. Speciation rates can vary; models of speciation rates include gradualism, in which speciation is slow and steady, and punctuated equilibrium, where speciation occurs in bursts followed by periods of stasis. Adaptive radiation is the formation of several new species from one common ancestor and occurs when new habitats become available Speciation can be rapid during times of ecological stress, such as the five major extinction events and current impact of humans on ecosystems Evidence supports the idea that evolution continues to occur, including chemical resistance, emergent diseases, and Darwin’s finches Explain the role of geographic isolation, reproductive isolation and polyploidy in the formation of new species Differentiate types of mating barriers Use data from a real or simulated population to predict future changes to the population Compare models of rates of speciation Describe speciation in an isolated population and connect speciation to microevolutionary forces such as selection and drift Evaluate the causes and effects of major extinction events, including human impact Design a plan for collecting data to investigate the claim that speciation and extinction have occurred Analyze data related to questions of speciation and extinction throughout Earth’s history Describe and evaluate data sets that illustrate ongoing evolution Evolution is always slow and steady Extinctions happened in the past Extinctions are always bad Populations are no longer evolving Natural processes can explain the origin of life. There are several hypothesis supported by scientific evidence about the origin of life on earth. The environment of primitive Earth contained inorganic molecules from which organic molecules could have been synthesized due to the presence of free energy and absence of oxygen Simple organic monomers could then serve as building blocks for more complex organic molecules, such as amino acids and nucleotides The joining of these polymers produced polymers with the ability to replicate, store and transfer genetic information The organic soup model hypothesize that these complex reactions could have occurred in solution; alternatively these reactions may have occurred on solid reactive surfaces The RNA World hypothesis posits that RNA could have been the earliest genetic material Geological evidence supports models of the origin of life on earth by providing a plausible range of dates for the origin of life Chemical experiments have shown that it is possible to form complex organic molecules from inorganic molecules in the absence of life Molecular and genetic evidence from existing and extinct organisms indicates that all organism on Earth share a common ancestor, include molecular building block common to all life forms and common genetic code Explain how the conditions of primitive earth provided an environment conducive to the evolution of the first cell Describe hypothesis for the origin of life, including the organic soup model and the RNA World hypothesis Evaluate scientific experiments and evidence that supports hypotheses for the origin of life Analyze molecular and genetic evidence that indicates all organisms on earth share a common ancestor Conduct and evaluate an experiment that simulates conditions for the origin of life on earth (Lab 4) Science cannot explain or provide evidence of the origin of life Essential Sub-Q Is evolution progress? Lesson Aim 1 2* Science Practice Given phenotypic data, propose an explanation for changes in phenotypic ratios by analyzing data sets and applying understanding of natural selection 1 – Models 5 – Data analysis 6 – Explanations Given a population of plants, investigate and explain how humans affect variation in other species by designing, conducting, and analyzing an artificial selection experiment 2 – Math 3 – Questions 4 – Data collection 5 – Data analysis 6 – Explanations Given case studies, identify and evaluate the effect of nonselective evolutionary forces on a population by explaining phenotype changes in various populations 1 – Models 5 – Data analysis 6 – Explanations Given quantitative data, apply the Hardy-Weinberg equation and conditions by calculating and explaining gene and genotypic frequencies in a population over time 2 – Math 4 – Data Collection5 – Data Analysis6 – Explanations Skill(s) Graphing Writing Text Analysis Inquiry Graphing Flip Learning Experiences Assessment Question Exam Alignment Natural selection Types of selection: directional, stabilizing, diversifying Rabbit/Wolf simulation Peppered moth activity Finch lab or case study In the case of peppered moths in industrial England, identify the type of selection that occurred, justify your conclusion with evidence and explain why this type of selection occurred PE #7-10, 37, IA #31, 2001 #2 Purpose of lab How to do lab Notebook set-up AP Lab 1 – Fast Plants Explain how the overuse of antibiotics has contributed to the current development of antibiotic-resistant strains of bacteria Genetic drift Gene Flow Mutation Nonrandom mating Cheating cheetahs Pom pom models Case Studies Identify and describe the evolutionary force that has imperiled the cheetah population, and discuss why cheating could save the species. PE # 3(II), 4G In 1950, the frequency of individuals with sickle cell anemia was 0.64. In 2010, the frequency of individuals with sickle cell anemia was 0.36. Determine the frequency of each allele in each generation and propose an explanation for the change in gene frequency. PE # 3(II), 4G HIV evolution PE # 3(II), 4G, 2008B #3 Populations can evolve, not individuals Investigative procedure for gathering data on natural selection Adaptations/Environment Distinguish correct definition of evolution from Lamarckian Identify graph that shows effect of darkened environment on phenotypes of beetles (directional) Writing Claims 4 5 Text analysis Diagram analysis Inquiry Graphing Writing Gene pools Calculating frequencies Deriving equation Example: Boobies Case studies Problem sets 7 – Connections Claims 6 Give data, apply the HardyWeinberg law and understanding of evolutionary forces by investigating, predicting, justifying the cause and effect of changes in the genetic makeup of a population (2 days 1 – Models 2 – Math 3 – Questions 4 – Data collection 5 – Data analysis 6 – Explanations 7 – Connections Text analysis Claims Writing How to use spreadsheet (Camtasia/Jing) Traditional lab with cards Spreadsheet scenarios Student-generated scenario Explain what terms in equation mean Describe conditions for equilibrium Calculate frequency of recessive allele if frequency of recessive trait is 12% Explain what terms in equation mean Describe conditions for equilibrium Calculate frequency of recessive allele if frequency of recessive trait is 12% Explain what terms in equation mean Describe conditions for equilibrium Calculate frequency of recessive allele if frequency of recessive trait is 12% How can we reconstruct evolutionary history? 7 8 9 Given data, justify the claim that all organisms share a common ancestor by describing and analyzing examples of conserved features and processes present in all domains of life 1- Models 5- Data analysis 6 – Explanations 7 - Connections Diagram analysis Writing Article analysis Prokaryotic vs. Eukaryotic cells 3 Domains Metabolic pathways “Junk” DNA Conserved vs. Derived Given diagrams and text, evaluate various types of evidence for evolution (including morphological, molecular, fossil and biogeographical) by inferring and justifying evolutionary relationships of groups 1 – Models 3 – Questions 5 – Data analysis 6 – Explanations Diagram analysis Article analysis Writing How to read a cladogram including homology hatch marks Types of evidence Radiometric dating Given morphological and molecular data, model the evolutionary relatedness of species by constructing and evaluating cladograms (2 days) 1 – Models 2 – Math 5 – Data analysis 6 – Explanations Claims Analyzing Diagrams Writing How to build a simple tree How to access and use BLAST 7 - Connections Tree vs. Circle of Life Cell models Journal articles? Case studies See BSCS Economist article on fossil vs. molecular evidence for human migration Cladogram practice Great Clad race (AMNH) AP BLAST Lab 3 Justify the claim that all organisms share a common ancestor using multiple pieces of evidence. A scientist has discovered 5 new species of bacteria in a deep-sea hydrothermal vent. Nucleotide sequences of each species are shown in the table above. Select the phylogenetic tree that is most consistent with the data, justify your conclusion and describe an additional source of evidence that could be used to improve or revise the Given the molecular data, build and justify a phylogenetic tree that illustrates the evolutionary relationship among these groups of organisms PE #5, 2004B #4, 1999 #3 All organisms share a genetic code organized into codons, an example of conserved processes PE #20, 58, 2011B #4b Similarities in protein structure suggest a common ancestor Method for finding and analyzing fossil evidence of extinction in various rock layers PE#32, 2009 #3, 2011B #4c Identify cladogram best supported by table of nucleotide differences How do new species evolve? 10 Given scenarios, apply the concepts of geographic isolation, reproductive isolation and polyploidy by proposing an explanation for the formation of two species 1 - Models 3 – Questions 5 – Data analysis 6 – Explanation 7 – Connections Article analysis Diagram Analysis Writing Mating barriers Allopatric Sympatric Polyploidy Mating barrier carousel Speciation simulation Polyploidy models Two populations of lizards have been found on opposite sides of a stream. The two populations are quite similar in appearance. Describe evidence that would confirm the hypothesis that these lizards are separate species. Explain how the two groups may have evolved into separate species. Connect your explanation to a force of PE # 4FR, 2011 #3c, 2011B #4a Describe two kinds of data that would show 2 groups are a species Explain how that data would show it (define biological species) microevolution 11 If all cells come from other cells, where did the first cell come from? 12 13 Given case studies, evaluate factors that affect rates of speciation by analyzing the cause and effect of a major extinction event 1 – Models 3- Questions 4 – Data analysis 6 – Explanations 7 – connections Text analysis Diagram analysis Writing Gradualism Punctuated equilibrium Adaptive radiation Fossil record analysis Mass extinction interactive Adaptive radiation case study (or other current example) Describe one major extinction event, identify factors that contributed to that event, and explain the impact of that event on species diversity 2004 #2, 2003B #4 Given scientific experiments, explain the origin of the first cell by describing and evaluating several hypotheses Including the organic soup model, heterotroph hypothesis and RNA World hypothesis 1 – Models 3 – Questions 5 – Data analysis 6 – Explanations Article Analysis Diagram analysis Anaerobic conditions on early earth Organic soup Protobiont Heterotroph hypothesis Endosymbiosis? Animations? Case study: Dating Earth Case study: Miller Case study: Oparin Summarize the experiment conducted by Stanley Miller, describe the hypothesis it was designed to test, and explain how its results support that hypothesis. PE #2, 8 (FR) Given experimental materials, investigate and explain conditions necessary for formation of cell membranes by designing, conduction and analyzing results of an experiment (2 days) 1 – Models 3 – Questions 4 – Data collection 5 – Data analysis Lab protocol Lab notebook Origin of Life lab Organic Soup Lab 7 – Connections 6 – Explanations Inquiry Claims Graphing Writing Journal - RNA vs. DNA Describe environmental conditions necessary for the formation of cell membranes. Identify purpose of Miller’s experiment as showing natural synthesis of organic molecules Identify evidence that would help answer question of whether microscopic organism was photosynthetic August Monday 20 Tuesday Wednesday Thursday Friday 21 22 23 24 Diagnostic Natural Selection Types of Selection Artificial Selection Lab 28 29 30 31 Hardy-Weinberg Hardy-Weinberg Lab Hardy-Weinberg Lab Quiz 3 4 5 6 7 NO SCHOOL Unity of Life Evidence for Evolution Cladograms Lab Cladograms Lab Course Intro, etc. 27 Microevolution Forces September 10 11 12 13 14 Speciation/Extinction lab Origin of Life Origin of Life Lab Origin of Life Lab 18 19 20 25 26 27 Speciation 17 21 Evolution Exam 24 28 AP Science Practices 1. 2. 3. 4. 5. 6. 7. Use representations and models to communicate scientific phenomena and solve scientific problems (Models) Use mathematics appropriately (Math) Pose, refine and evaluate scientific questions (Questions) Plan and implement data collection strategies appropriate to a particular scientific question (Data collection) Perform data analysis and evaluation of evidence (Data Analysis) Construct, justify, refine, and evaluate scientific explanations and theories (Explanations) Connect and relate knowledge across various scales, concepts and representations in and across domains (Connections)