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Kimberly Pambid 1st period AP BIO 10/02/09 FREE RESPONSE A fixed action pattern (FAP) is an instinctive behavioral response that is triggered by a very specific stimulus. Fixed action patterns are produced by a neural network known as the innate releasing mechanism. The FAP is triggered in response to an external sensory stimulus known in animal behavior terms as a sign stimulus, or, if it is a signal from one individual to another, it is called a releaser. Tinbergen envisioned such a mechanism for each of the major functional categories of behavior: foraging, reproduction, anti-predator behavior, etc. Nikolaas "Niko" Tinbergen was a Dutch ethologist who shared the 1973 Nobel Prize in Physiology with Karl von Frisch and Konrad Lorenz for their discoveries concerning organization and elicitation of individual and social behavior patterns in animals. The example that Tinbergen worked out in greatest detail is the reproductive instinct of the fish known as the three-spined stickleback. The male turns a bright red/blue color during the mating season. This color change is the FAP in response to an increasing day length which is the sign stimulus. During this time they are also naturally aggressive towards other red-bellied sticklebacks, another FAP. However anything that is red, or has the appearance of being red, will bring about this FAP. The proximate response to this is that due to the stimuli, a nerve sends a signal to attack that red item. The ultimate cause of this behavior stems from the fact that the stickleback needs the area in which it is living for either habitat, food, mating with other sticklebacks, or other purposes. Imprinting is the term used in psychology and ethology to describe any kind of phase-sensitive learning (learning occurring at a particular age or a particular life stage) that is rapid and apparently independent of the consequences of behavior. It was first used to describe situations in which an animal or person learns the characteristics of some stimulus, which is therefore said to be "imprinted" onto the subject. The best known form of imprinting is filial imprinting, in which a young animal learns the characteristics of its parent. It was rediscovered by the early ethologist Oskar Heinroth, and studied extensively and popularized by his disciple Konrad Lorenz working with graylag geese. Lorenz demonstrated how incubator-hatched geese would imprint on the first suitable moving stimulus they saw within what he called a "critical period" between 13–16 hours shortly after hatching. Most famously, the goslings would imprint on Lorenz himself, and he is often depicted being followed by a gaggle of geese who had imprinted on him. Filial imprinting is not restricted to animals that are able to follow their parents, however; in child development the term is used to refer to the process by which a baby learns who its mother and father are. The process is recognized as beginning in the womb, when the unborn baby starts to recognize its parents' voices. Another one would be sexual imprinting, which is the process by which a young animal learns the characteristics of a desirable mate. Classical conditioning (also Pavlovian or respondent conditioning) is a form of associative learning that was first demonstrated by Ivan Pavlov, a Russian physiologist who loved to work with dogs and their digestive process. The typical procedure for inducing classical conditioning involves presentations of a neutral stimulus along with a stimulus of some significance. The neutral stimulus could be any event that does not result in an overt behavioral response from the organism under investigation. Pavlov referred to this as a conditioned stimulus. Conversely, presentation of the significant stimulus necessarily evokes an innate, often reflexive, response. Pavlov called these the unconditioned stimulus and unconditioned response, respectively. The original and most famous example of classical conditioning involved the salivary conditioning of Pavlov's dogs. During his research on the physiology of digestion in dogs, Pavlov noticed that, rather than simply salivating in the presence of meat powder, the dogs began to salivate. Pavlov called these psychic secretions. From this, he predicted that, if a particular stimulus in the dog’s surroundings were present when the dog was presented with meat powder, then this stimulus would become associated with food and cause salivation on its own. In his initial experiment, Pavlov used a bell to call the dogs to their food and, after a few repetitions, the dogs started to salivate in response to the bell. Thus, a neutral stimulus (metronome/bell) became a conditioned stimulus as a result of consistent pairing with the unconditioned stimulus. In 1947, Karl von Frisch, an Austrian ethologist who received the Nobel Prize in Physiology or Medicine in 1973, along with Nikolaas Tinbergen and Konrad Lorenz, correlated the runs and turns of the bee dance to the distance and direction of the food source from the hive. The orientation of the dance correlates to the relative position of the sun to the food source, and the length of the waggle portion of the run is correlated to the distance from the hive. Also, the more vigorous the display is, the better the food. There is no evidence that this form of communication depends on individual learning. One of the most important lines of evidence on the origin and utility of the dance is that all of the known species and races of honey bees exhibit the behavior, but details of its execution vary among the different species. For example the "dwarf honeybees" dance is performed on the dorsal, horizontal portion of the nest, which is exposed. The runs and dances point directly toward the resource in these species. Each honey bee species has a different correlation of "waggling" to distance, as well. Such species-specific behavior suggests that this form of communication does not depend on learning but is rather determined genetically. It also suggests how the dance may have evolved. Frisch's honey bee work included the study of the pheromones that are emitted by the Queen bee and her daughters, which maintain the hive's very complex social order. Pheromones are chemical odors one organism emits to communicate with another organism, most likely for reproductive reasons. Outside the hive, the pheromones cause the male bees, or drones, to become attracted to a queen and mate with it. Inside the hive, the drones are not affected by the odor. Operant conditioning is the use of consequences to modify the occurrence and form of behavior. Operant conditioning is distinguished from classical conditioning in that operant conditioning deals with the modification of "voluntary behavior" or operant behavior. Operant behavior "operates" on the environment and is maintained by its consequences, while classical conditioning deals with the conditioning of respondent behaviors which are elicited by antecedent conditions. Behaviors conditioned via a classical conditioning procedure are not maintained by consequences. Burrhus Frederic Skinner was an American Psychologist, author, inventor, advocate for social reform, and poet. Skinner conducted pioneering research and created his own school of Radical Behaviorism, which seeks to understand behavior as a function of environmental histories of reinforcing consequences. He is known as the inventor of the operant conditioning chamber (or Skinner box), a research tool used to examine the orderly relations of the behavior of organisms (such as rats, pigeons and humans) to their environment. He discovered what is now called operant conditioning and articulated the now widely accepted term reinforcement as a scientific principle of behavior. Skinner understood the development of behavior as occurring through the effects of positive and negative reinforcement as well as punishment and extinction. Reinforcement processes were emphasized by Skinner, and were seen as primary in the shaping of behavior. A common misconception is that negative reinforcement is some form of punishment. Punishment and extinction have the effect of weakening behavior, or decreasing the probability of a behavior reoccurring, by the application of an aversive event (punishment) or the removal of a rewarding event (extinction). For example, a box where a pigeon or rat had to press a button in order to receive food. Skinner found that the pigeon/rat eventually would stumble upon the fact that pushing the button elicited a food pellet. The pigeon would become conditioned to press the button in order to receive the pellet, and the frequency of the conditioned response--pushing the button--would increase. At the same time, there would also be buttons that would shock them and as they keep doing it, they would eventually learn to stop because it hurts them.