* Your assessment is very important for improving the workof artificial intelligence, which forms the content of this project
Download Ecology Introduction 1. Ecology
Conservation agriculture wikipedia , lookup
Photosynthesis wikipedia , lookup
History of wildlife tracking technology wikipedia , lookup
Triclocarban wikipedia , lookup
Sustainable agriculture wikipedia , lookup
Theoretical ecology wikipedia , lookup
Lake ecosystem wikipedia , lookup
Aftermath: Population Zero wikipedia , lookup
Renewable resource wikipedia , lookup
Ecology Introduction Ecology is the study of the natural environment and organisms, how they interact and how they change their environment. There is a dynamic relationship between every organism in any environment, as well as between the organisms and the physical environment. For conservation to be successful in Namibia, it is essential that every inhabitant is familiar with the basic principles of ecology. The aim of this unit on ecology is not to teach you about conservation. The emphasis in this unit will be to trace the energy pathway from the sun through living things and identify the roles of various organisms (producers, consumers, decomposers, predators, prey) in a food chain and a food web. Upon completion of this unit you will be able to: explain the essentials of ecology, some ecological components, principles and terminology; explain how all factors, living and non-living, are dependent on each other and are in constant interaction with one another explain what an ecosystem is, and how we can study one explain the application of ecological concepts into everyday living summarise the concept of energy flow through a food web 1. Ecology 1.1 Origin of the word…”ecology” The word ecology has a Greek origin OIKOS = household LOGOS = study of… Study of the “house/environment” in which we live. The study of the interactions that take place among organisms and their environment 1.2 What is an environment? Everything that affects an animal and plant makes up its environment - where it lives, the weather and all the living things it comes into contact with. Every living thing, including people, has an effect upon the environment. 1.3 Animal and Plant Adaptations All living things have to be suited to their environment if they are to survive. Fish have streamlined bodies, fins and specially shaped tails to help them move quickly and easily through the water. Ground squirrels have bushy tails to shade them from the hot Namibia sun. Squirrels have sharp, strong claws for gripping tree trunks and branches and strong teeth for eating nuts. 1.4 What is a habitat? The place in which an organism lives provides the kinds of food and shelter, the temperature, and the amount of moisture the organism needs to survive. 194 Alternatively, you can also say that a habitat is any place where organisms live together naturally. It is like a neighbourhood in nature. 1.5 What does an organism need in a habitat? In a habitat an organism need shelter, water, food and plants need sunlight (Fig 45). Figure 45: Shelter Water Food Difference between environment and habitat Environment – condition in which an organism lives e.g. temperature, light, humidity etc. these be affected by other organisms. Habitat – specific environment in which organisms’ lives. 1.6 What about our habitat? Just like other animals and plants, we need our own habitat. What are our needs? Food, shelter and water (fig 46) Figure 46: Food Shelter 195 Water What is the difference between environment and habitat? Environment includes all living & non-living things that exist on this earth. The living things (biotic factors) include all the plants, animals including man, & micro-organisms. The non-living factors (abiotic factors) include the air, light, water, temperature, soil, humidity, rainfall etc. A habitat is a specific natural environment. The area or environment where an organism or ecological community normally lives or occurs: a marine habitat. 1.7 Biosphere The part of Earth that supports life is the top portion of Earth's crust, all the waters that cover Earth's surface, and the atmosphere that surrounds Earth. The biosphere, like the human body, is made up of systems that interact and are dependent on each other. 1.8 Ecosystem The biosphere’s systems are called Ecosystems. All ecosystems must have a constant source of energy (usually the sun) and cycles or systems to reuse raw materials. Examples are the water, nitrogen and carbon cycles. An ecosystem is made up of all the biotic or living and the abiotic or non-living components in a given area. The study of ecosystems mainly consists of the study of certain processes that link the living or biotic components to the non-living or abiotic components 1.9 What is a Population? All the members of a species living in one place at one time. Example – All the terrapin in Avis Dam or all the barbel (catfish) in Avis dam or all the sweethorn trees growing around Avis Dam. Figure 47: Terrapin (left). Pelicans (right) Source: B. Curtis 1.10 What is a community? All the species in a given area (or) all populations (different species) that live in a particular area. Example – all the living things in Goreangab dam or Avis dam. In Goreangab dam we find – terrapin – barbels – frogs – pelicans etc. 196 Figure 48: The community of animals along the Chobe River consists of a population of waterbuck and a population of baboons (as well as other animals and plants). Source: B. Curtis Density and dispersion patterns are important population variables Two important aspects of population structure are population density and dispersion pattern. Population density is the number of individuals of a species per unit area or volume-the number of oak trees per square kilometre (km 2) in a forest, for instance, or the number of earthworms per cubic meter (m3) in forest soil. Because it is impractical or impossible to count all individuals in a population in most cases, ecologists use a variety of sampling techniques to estimate population densities. For example, they might base an estimate of the density of alligators in the Florida Everglades on a count of individuals in a few sample ploys of 1 km2 each. The larger the number and size of sample plots, the more accurate the estimates. In some cases, population densities are estimated not by counts of organisms but by indirect indicators, such as number of bird nests or rodent burrows. Within a population’s geographic range, local densities may vary greatly. The dispersion pattern of a population refers to the way individuals are spaced within their area. A clumped dispersion pattern, in which individuals are grouped in patches, is the most common in nature. Clumping often results from an unequal distribution of resources in the environment. For instance, plants or fungi may be clumped in areas where soil conditions and other factors favour germination and growth. Clumping of animals is often associated with uneven food distribution or with mating or other social behaviour. For example, fish are often clumped in schools, which may reduce predation risks and increase feeding efficiency. 197 A uniform dispersion pattern (an even one) often results from interactions between the individuals of a population. For instance, some plants secrete chemicals that inhibit the germination and growth of nearby plants that could compete for resources. Animals may exhibit uniform dispersion as a result of territorial behaviour shows uniform dispersion in king penguins. 198 In a random dispersion pattern, individuals in a population are spaced in an unpredictable way, actually without pattern. Plants, such as dandelions, that grow from windblown seeds might be randomly dispersed. However, varying habitat conditions and social interactions make random dispersion rare. Estimates of population density and dispersion patterns enable researchers to monitor changes in a population and to compare and contrast the growth and stability of populations in different areas. 199 1.11 Abiotic factors The nonliving things in an environment are called abiotic factors. Examples of abiotic factors are sunlight, temperature, water, rainfall, climate and soil (fig 49). Figure 49: clouds are examples of abiotic factors. 1.11 The abiotic components 1.11.1 Topography Topography is important since it includes such factors as the height above sea level and the slope and degree of exposure by sunlight. North-facing slopes receive more sunshine and the soil temperature is higher. The water content of the soil is lower, while that of the south-facing slopes is higher. The soil is cooler and there are also more plants on the south-facing slopes. 1.11.2 Edaphic or soil factors Soil is obtained through the weathering of rocks by physical, chemical and biotic processes. 200 Soil is classified on the basis of soil particle size: Eg; sand, clay and loam. Humus makes soil more fertile, assists in binding sandy soil and increases the water-holding capacity of the soil. The depth of soil is very important. The upper layer is formed continuously over many years –about 30mm over 500 years! The type and the composition of the soil influences the biotic components, but the biotic components also influence the soil. Example: the enrichment of the soil and the formation of humus during the decomposition process of biotic components. 1.11.3 Climatic factors 1 Light Plants and animals are adapted to light in many ways, including the following: (i) Light is essential for plants because it is needed for photosynthesis. (ii) Animals such as the earthworms, bats, and lions are active at night. 2 Temperature Temperature influences the biological functioning of organisms and therefore their distribution around the world. Some animals are cold-blooded, for example fish, amphibians and reptiles. Their temperature varies according to the environment’s temperature. When it is cold their body metabolism activity decreases (hibernate). Other animals are warm-blooded, for example mammals and birds. They maintain a constant body temperature which is not affected by external temperature changes. Some warm-blooded animals hibernate during the cold season (bats). Others migrate to warmer areas and return in the spring (swallow, storks). Temperature influences plant life activity E.g. fruit trees and plants such as grapes lose their leaves in winter and thereby limit their metabolism. Hibernate – to pass the winter in a dormant condition with metabolism greatly slowed down. 1.11.4 Water and the water cycle a) Water is probably the most important ecological factor. It is responsible for all physical processes which occur in plant and animals. b) Once water has been used, it does not disappear. Instead, it is recycled. c) The water cycle plays a very important role in plant and animal life: The water cycle is the continuous movement of water in a cycle between the soil and the atmosphere. d) Referring to figure 50, the following occurs during the water cycle: 201 Figure 50: The water cycle i) Precipitation: frost, mist, hail, snow, and rain: this is the water moving from the atmosphere to the soil, providing moisture, absorbing the heat and cooling the soil. ii) The water that has fallen is absorbed by the upper layers of soil. Infiltration: the water moves through the topsoil into the subsoil and into the water table where it is stored, and becomes available for plant roots. Some of the water also flows away into the dams and the sea. iii) Evaporation: the water reaches the atmosphere again from the large water surfaces that make up 70% of the earth’s surface, and also by means of evaporation from the soil. iv) Transpiration: plants lose water by means of transpiration and this reaches the atmosphere. 1.11.5 Fires Fire is a natural part of the sawannah ecosystems in many parts of the world e.g. Namibia. It can be good for plants. In some plants the germination of seed is stimulated by fire. However, too much fire or fire at the wrong time of the year is extremely bad to the environment. Uncontrolled and unplanned fires cause a lot of damage ecologically and economically and must be prevented. 1.11.6 Atmospheric gases All living organisms are dependent on the air around them. The most important gases for animals and plants are: oxygen, carbon dioxide, nitrogen and water vapours. In the atmosphere there is about 21% oxygen, 0.03% C02 and 79% Nitrogen. Oxygen is obtained from photosynthesising plants. Carbon dioxide in the atmosphere comes from the carbon dioxide in the soil and in the water. It is also released during the decomposition of organic material and as a by-product of respiration (breathed out by humans and animals) Nitrogen is important for the synthesis of substances such as proteins. It is produced by Nitrogen-fixing bacteria in the soil and is also made available to plants in the form of nitrates during thunderstorms. 202 All the elements essential for life move in cycles. Without this cycling of nutrients and elements, the natural resources would soon be exhausted. Two such cycles, namely the oxygen and the carbon dioxide cycles will be discussed. 1. The oxygen cycle Atmospheric oxygen is important for the cellular respiration in living organisms. During respiration, oxygen combines with hydrogen, and water is formed in the cells. This water can be excreted by the organisms. It is also used by green plants for photosynthesis. A by-product of photosynthesis is oxygen-and this oxygen is made available for respiration once again. Atmospheric oxygen in the form of O is part of the ozone layer which helps to prevent highenergy radiation from the sun reaching the earth. 2. The carbon dioxide (carbon) cycle Carbon dioxide is built up from carbon and oxygen. Carbon is essential for all living organisms. Carbon dioxide is constantly being produced: 1) By the respiration processes of animals on land and in the sea; ii) Through the burning (combustion) of oil, petrol, coal, by motorcars, factories, aeroplanes and households; iii) Through fires (veld fires, household fires); iv) Bacterial respiration during decomposition: at the death of a plant or an animal. These plant and animal remains will during the course of millions of years again form products such as oil and coal. Removal of carbon dioxide occurs through the photosynthetic processes of plants. During photosynthesis, the carbon dioxide from the atmosphere is used to make carbon products such as glucose. 1.12 Biotic factors Biotic factors are all the living things or their materials that directly or indirectly affect an organism in its environment. Factors such as parasitism, disease, and predation (one animal eating another) would also be classified as biotic factors. 1.13 The biotic components All the living factors in an ecosystem are known as the biotic components. The biotic components of an ecosystem can be divided into two large groups 1 the autotrophic component (self feeding – make their own food) 2 the heterotrophic component (various feeding) 203 1.13.1 The autotrophic components Organisms that can use the energy in sunlight to convert water and carbon dioxide into Glucose (food - photosynthesis) Autotrophs are also called Producers because they produce all of the food that heterotrophs use Without autotrophs, there would be no life on this planet E.g. Plants and Algae Figure 51: Autotrophs that get their energy from inorganic substances, such as salt are called Chemotrophs. They live deep down in the ocean where there is no sunlight. E.g. Bacteria and Deep Sea Worms 1.13.2 The Heterotrophic components Organisms that do not make their own food Another term for heterotroph is consumer because they consume other organisms in order to live. Heterotrophs live from the organic substances produced by the autotrophs – the photosynthesizing green plants E.g. Rabbits, antelope, mushrooms Figure 52: Impala 204 Source: B. Curtis Heterotrophs may be divided into the following groups: 1 The primary consumers These are also known as the herbivores. They are plant-eaters and live on the producers or the green plants Examples: garden snails, caterpillars, locusts, mice, giraffe, cows, elephants, hippos, rhinos, antelope. Figure 53: Giraffe Source: B. Curtis The secondary consumers Carnivores – eat ONLY meat. They include the predators. They prey on the herbivores and other (smaller) carnivores. E.g. lions, lizards, spiders, leopard, cheetah,sharks Figure 54: Cheetah (left). Leopard (right) 205 Source: B. Curtis Predator A predator eats other animals. For example: The cat is a predator, because it eats other animals. The bear is a predator, because it eats other animals. Figure 55: Cats (left) and bears (right) eat fish People are predators too! Figure 56: Prey Any animal which is hunted and killed by another animal for food is prey. Figure 57: Predator Prey 206 3. The omnivores Omnivores – eat BOTH plants and animals. Omnivores are therefore also heterotrophs and may be primary or secondary consumers. Eg; human beings, pigs, crabs, baboons Figure 58: Human being (left) Source: humanitarianreform.org 4. Scavengers/Detritivores – feed on the tissue of dead organisms (both plants and animals) or they eat the remains of dead organisms left by the consumers Eg. – Vultures, hyaenas, crows and shrimp Figure 59: Black-backed jackals (left) Vultures (right) Source: B. Curtis 1.13.3. The decomposers The last links in the chain are the decomposers. They are also known as the microconsumers. They are also heterotrophs which feed on decaying or dead matter. (They break down all dead organic matter). If you defecate (poop), they eat that. If you lose a leaf, they eat it. 207 Whenever something that was alive dies, the decomposers get it. Decomposers break down nutrients in the dead material and return it to the soil. The producers can then use the nutrients and elements once they are in the soil. The decomposers complete the system, returning essential molecules to the producers. Decomposers are very important since they break down complex organic substances into simpler substances which are released into the environment and absorbed by the producers. E.g. – Bacteria, insect larvae and Mushrooms 1.14 Trophic levels Energy moves from one organism to another when food containing the energy. Each step in this transfer of energy is known as a trophic level. The main trophic levels are producers, consumers, and decomposers. There is a constant flow of energy between these trophic levels and a contanst loss from one level to the next. The primary consumers obtain their energy from the producers; the secondary consumers eat the primary consumers and/or other secondary consumers and in this way obtain their energy. The cycle ends when the producers and consumers are decomposed by the decomposers to form abiotic components again. Decomposers – bacteria and fungi that break down dead organisms and recycle the material back into the environment. There is therefore a constant flow of energy from the sun through different organisms and back to the soil and atmosphere. 1.15 Food chains The energy flow from one trophic level to the other is known as a food chain. A food chain is simple and direct. It involves one organism at each trophic level. This transfer or flow of energy from its source (the sun) and then through green plants and a series of other organisms – each of which is food for the next – is known as a food chain. Figure 60: 208 1.16 Food web Most organisms eat more than JUST one kind of organism. When more organisms are involved it is known as a FOOD WEB. Food webs are more complex and involve lots of organisms. Isolated single food chains as shown in figure 58 never occur in nature. Usually a number of food chains are interwoven to form a large and complex food web. Food webs show what eats what in order to gain the energy it needs to live. Food webs use arrows to show what eats what. Nearly all food webs start with a green plant. Green plants are called producers because they produce their own food. Animals are consumers. Some consume plants to get the energy they need to live (primary consumer). Others consume other animals (secondary consumers – they eat the primary consumers). Animals that are eaten by other animals are called prey . Some prey animals are herbivores. Some omnivores and carnivores are also prey to other animals. Animals that eat other animals are called predators. Some animals are predators and prey. This means they eat some animals but are eaten by others. The top predator that is prey to no other animal is called an apex predator. Eg; lion. However if a lion dies it is eaten by scavengers (Eg; vultures and jackal) and decomposed by bacteria. Figure 61: Food web 209 1.17 Ecological pyramid An ecological pyramid shows the relationship between producers and consumers at different trophic levels in an ecosystem. It shows the relative amounts of energy or matter contained at each trophic level. The Pyramid shows which level has the most energy and the highest number of organisms. The producers form the base of the pyramid (fig 60) and the consecutively smaller trophic levels are placed higher up the pyramid until the apex is reached. With these pyramids it can be seen that the producers are the most numerous, the secondary consumers still fewer, and so on. There are three types of pyramid that can be used to display ecological systems: Pyramids of NUMBER Pyramids of BIOMASS Pyramids of ENERGY Figure 62: Which level has the most energy? Which level has the most organisms? Which level has the least organisms? Which level has the least energy? 1 Pyramid of Numbers The food chains and webs show the flow of material and energy in ecosystems, but they do not give you any idea of how many organisms there are in the ecosystem. A pyramid of numbers (fig. 61) shows the numbers of organisms present in each level of a food chain. Suppose the numbers found in a particular habitat are as follows: 2000 plants 25 mice 1 owl The pyramid of numbers would be as shown below. 210 Figure 63: Pyramid of Numbers In some cases, however, a numbers pyramid can seem inverted. An inversion seems to exit when an organism that occurs in the greater number does not fall at the bottom of the pyramid. Sometimes you do not get a pyramid because one organism in the chain is unusually large, e.g. one large tree providing food for thousands of caterpillars and hundreds of secondary consumers, such as spiders and birds (fig 64). Figure 64: numbers pyramid can seem inverted 2 Pyramids of Biomass 211 Biomass is a word used to describe the mass of living material in an area. If you could collect all of the plants and animals in an area and weigh them, this would be the biomass of that area. A pyramid of biomass is a diagram showing the total weight (mass) of organisms at each level of a food chain. Biomass always gives a pyramid since producers must have the most biomass and consumers must have less. Measured in Kg, grams. Pyramids of biomass are always pyramid-shaped. Figure 65: Pyramids of Biomass 3 Pyramid of Energy Shows the energy available at each trophic level. The size of the blocks represents the proportion of productivity. Measured in Joules or Calories. Most of the energy available to the community is in the 1st trophic level. Only 10-20% of the energy is available to the next trophic level (≈ 90% lost). 212 Figure 66: Pyramid of Energy This is the most important pyramid since energy is what is really being transferred from one level to the next. Just as numbers of plants or animals can be misleading (fig 66) so can biomass e.g. 1kg of grass contains less energy than 1 kg meat. 1.18 Breaking the chain Organisms living in a ecosystem depend on each other. If one part of a food chain dies out or is greatly reduced, the consumers have to find alternative food, move away, or starve. This then affects the consumers further along the chain/web. 1.19 Endangered or extinct? The number of people in the world is growing at an alarming rate. But this is not true for all animals and plants. In some cases, there are only a few of one type of animal and plant left in the wild. These organsiums are endangered. If they die out completely, they become extinct. 1.20 Why does this happen? There are lots of reasons why organsims become endangered or extinct. The most common are: Loss of habitat (woodlands cut down, rivers drying up, bush encroachment). Chemicals or pollution poison the animals. Overexploitation – too much use e.g. hunting (for sport, their fur, tusks or meat) or collecting (for firewood etc.) 1.21 Caring for the environment It is in our own best interests to look after the world we live in. If a habitat is lost or damaged, it has an effect on everything else, even if we do not see or understand it straight away. Remember - once something becomes extinct, it’s gone forever! We can mismanage the environment and cause it to become unproductive e.g. too many herbivores on a farm will kill the grass and lead to erosion. 213 1.22 The mutual interactions in an ecosystem As you must have seen during walks through the bush and the veld, a number of interesting interactions occur in nature, ensuring the survival of a particular species: shrubs provide twigs for spiders to spin their webs insects use leaves on which to lay their eggs plants depend on animals for the dispersal of seed large trees offer shade to plants and to animals There is an endless list of these interactions occurring in nature. Let us study these interactions, as well as the different forms of interactions, in some more detail: 1.23 Competition Organisms within a community often have to share water, food and living space. When there is enough of everything for everybody, then sharing is easy. However, if one factor is limited (scarce), the organisms will compete with one another. Example: (i) competition between animals for a limited supply of food, water or shelter (ii) Competition between plants to obtain enough sunlight or space to grow, example seeds which have been sown too close together. Intraspecific competition is competition between two or more individuals of the same species, because their needs are similar. E.g. lions fighting at a kill Interspecific competition is competition between organisms of different species (ex. between kudu and giraffe) 1.25 Symbiosis A close and permanent association between organisms of different species. 1 Commensalism – a relationship in which one organism benefits and the other is not affected Example: cow seeking shade beneath a tree Bird nest in the tree Bromiliad Roots on Tree Trunk Without Harming Tree 214 2 Mutualism – a relationship in which both organisms benefit from each other Example: Birds eating pest off a rhino’s back Bees visit a flower to obtain nectar; the flowers need the bee for pollination. Anemones Protect and Feed Clownfish 3 Parasitism – A relationship in which one organism benefits and the other is harmed Example: Ticks on a dog Tapeworm (Taenia solium) lives in the intestine of man 215 There are two types of parasites: a) The ectoparasite living outside the body of the host: - ticks (parasite) on cattle (host). - lice on fowls; - -fleas on dogs and cats - Aphids on plants b) The endoparasites living inside the body of the host; in the digestive system, in the blood vessels, the tissues, the air passages: 4 - bilharzias (schistosoma) lives in the bladder or the intestine of man - the plant parasite: dodder, living on Lucerne, sunflowers and mealies. Predation- Predators are secondary consumers and eat other living animals Predators have a number of victims (parasites normally have only one). They either eat their entire prey or leave the rest to scavengers Example: lions, hawks, snakes, parying mantis, spiders. Disturbances Disturbances are events such as storms, fire, floods, droughts, overgrazing, or human activity that damage biological communities, remove organisms from them, and alter the availability of resources. The types of disturbances and their frequency and severity vary from community to community. By gathering data from specific communities over many years, ecologists are beginning to appreciate and understand the impact of disturbances. We think disturbances in negative terms, but not always. Small-scale disturbances often have positive effects. For example, when a large tree falls in a windstorm, it disturbs the immediate surroundings, but it also creates new habitats. For instance, more light may now reach the forest floor, giving small seedlings the opportunity to grow; or the depression left by its roots may fill with water and be used as egg-laying sites by frogs, salamanders, and numerous insects. Small-scale disturbances may enhance environmental patchiness, which can contribute to species diversity in a community. Communities change drastically following a severe disturbance that strips away vegetation and even soil. The disturbed area may be colonized by a variety of species, which are gradually replaced by a succession of other species, in a process called ecological succession. 216 Ecological Succession • • A change in the community in which new populations of organisms gradually replace existing ones – 1. Primary Succession – occurs in an area where there is no existing communities and for some reason (s) a new community of organisms move into the area A change in the community in which new populations of organisms gradually replace existing ones – 2. Secondary Succession – occurs in an area where an existing community is partially damaged – 3. Climax Community – a community that is stable and has a great diversity of organisms An example of succession 1. There are certain grasses, example the Aristida, which are known as pioneer grasses. As the grass grows and dies, humus and soil is formed. Insects and decay microbes are also part of this community. 2. This pioneer community changes the abiotic surrounding and creates favourable conditions for other organisms: secondary succession can now take place. Shrubs and bushes grow, mice, jackals and birds arrive. 3. Next, small trees, then taller trees grow. The variety and number of plant species increases. 4. Eventually a forest with a mature, self-supporting climax community is formed. 5. Climax communities may be formed in the veld, a dam, or in the ocean. 217