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Evolution of Biological Communities Outline I. Evolution by Natural Selection A. Background B. Elements of Theory C. Examples II. Population Genetics Background Charles Darwin was not the first scientist to think about life’s origins or about how species might change over time. Lamarck – – Study of fossils suggested life evolves Inheritance of acquired characteristics Charles Darwin (1809-1882) Even as a boy, interested in nature Accumulated observations leading to theory – – – – Lyell’s geology Galapagos Islands Artificial breeding programs Malthus’ theories of population growth Through these observations, Darwin began to see adaptation and the origin of new species as closely related processes Descent with modification Evolution by Natural Selection Observation: In any species, more offspring are born into a population than actually survive to reproduce Observation: Resources in any environment are limited Inference: production of more individuals than the environment can support leads to a struggle for existence between individuals of a population, with only a fraction of their offspring surviving each generation Evolution by Natural Selection Observation: Members of a population vary in their characteristics Observation: Much of this variation is heritable Inference: Survival depends in part on inherited traits. Individuals whose inherited traits give them a high probability of surviving and reproducing will leave more offspring than those without these inherited traits Inference: leads to gradual accumulation of these favorable characteristics in the population over generations Natural Selection Adaptations: traits which give a selective advantage to some individuals in a population ie. traits which increase an organisms chances of survival and reproduction Selective Agent/Pressure: environmental factors that influence which individuals leave more offspring (favor certain adaptations within the population) Natural Selection is the name given to the process of differential reproduction-the mechanism by which evolution takes place Important Points Being “fit” is relative, if the environment changes, new traits may be favored and individuals suited to one environment may not be suited anymore It is not “survival of the fittest” but survival and reproduction of the fittest! Natural selection does not have foresight-no direction toward the “perfect” organism-blind process whereby existing variation is sorted out by fitness. Population Genetics Variation – – – Heredity an important and necessary part of the theory, not well understood in Darwin’s time As study of heredity and genetics developed, the underlying control of an organisms traits has become better understood 2 types of variation Non-heritable: no genetic basis, can’t be passed down, often of environmental origin Heritable: actual change in the DNA of an organism via mutation or mutagen, passed down via egg or sperm Non-heritable Variation Heritable Variation Population Genetics Gene: specific sequence of nucleotides in DNA which code for a particular protein Allele: alternate forms of a gene – Examples More precise definition of evolution: Evolution is a change in the allele frequencies of a population of organisms over time Population: a group of interbreeding individuals, live in the same area at the same time