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Ecosystems Chapter 26 LEARNING OBJECTIVE 1 • Define ecology • Distinguish among population, community, ecosystem, and biosphere KEY TERMS • ECOLOGY • • Discipline of biology that studies interrelations between living things and their environments Ecologists study populations, communities, ecosystems, and the biosphere KEY TERMS • POPULATION • • A group of organisms of the same species that live in a defined geographic area at the same time COMMUNITY • An association of populations of different species living in a defined habitat with some degree of interdependence Community: A Nurse Log KEY TERMS • ECOSYSTEM • • Interacting system that encompasses a community and its nonliving, physical environment BIOSPHERE • All of Earth’s living organisms, collectively LEARNING OBJECTIVE 2 • Explain the difference between Jshaped and S-shaped population growth curves Dispersion in Populations (a) Random dispersion (b) Clumped dispersion (c) Uniform dispersion Fig. 26-2, p. 517 Intrinsic Rate of Increase • The maximum rate at which a population could increase in number under ideal conditions Exponential Population Growth • J-shaped curve • An accelerated pattern of growth exhibited by certain populations for a limited period Exponential Population Growth Logistic Population Growth • S-shaped curve • After accelerated growth, growth rate decreases • Natural populations seldom follow the logistic growth curve closely Logistic Population Growth and Carrying Capacity LEARNING OBJECTIVE 3 • Summarize the three main types of survivorship curves Survivorship • The probability that a given individual in a population will survive to a particular age 3 General Survivorship Curves • Type I survivorship • • Type II survivorship • • Mortality greatest in old age Mortality spread evenly across all age groups Type III survivorship • Mortality greatest among the young Survivorship Curves Experiment: Drummond Phlox Metapopulation LEARNING OBJECTIVE 4 • Characterize producers, consumers, and decomposers KEY TERMS • PRODUCER • An organism that synthesizes organic compounds from simple inorganic raw materials KEY TERMS • CONSUMER • • An organism that cannot synthesize its own food from inorganic raw materials Must obtain energy and body-building materials from other organisms KEY TERMS • DECOMPOSER • A microorganism that breaks down dead organic material and uses the decomposition products as a source of energy LEARNING OBJECTIVE 5 • Describe what is meant by an organism’s ecological niche KEY TERMS • ECOLOGICAL NICHE • The totality of an organism’s adaptations, its use of resources, its interactions with other organisms, and the lifestyle to which it is fitted in its community LEARNING OBJECTIVE 6 • Define competition, predation, and symbiosis • Distinguish among mutualism, commensalism, and parasitism KEY TERMS • COMPETITION • Interaction among two or more individuals that attempt to use the same essential resource, such as food, water, sunlight, or living space KEY TERMS • PREDATION • A relationship in which one organism (the predator) kills and devours another organism (the prey) KEY TERMS • SYMBIOSIS • • An intimate relationship between two or more organisms of different species Symbiosis includes mutualism, commensalism, and parasitism Symbiosis • Mutualism • • Commensalism • • • Both partners benefit One organism benefits Other is unaffected Parasitism • • One organism (the parasite) benefits Other (the host) is harmed Plant Defense Against Herbivores Mutualism Commensalism LEARNING OBJECTIVE 7 • Define ecological succession • Distinguish between primary succession and secondary succession KEY TERMS • ECOLOGICAL SUCCESSION • Sequence of changes in the species composition of a community over time Succession • Primary succession begins in an area not previously inhabited • • Example: bare rock Secondary succession begins in an area where there was a preexisting community and a well-formed soil • Example: abandoned farmland Primary Succession Lichens Exposed and mosses rocks Ferns, grasses, and herbs Low shrubs Jack pine, black spruce, and aspen Balsam fir, paper birch, and white spruce forest community Fig. 26-12, p. 526 Secondary Succession LEARNING OBJECTIVE 8 • Summarize the concept of energy flow through a food web KEY TERMS • TROPHIC LEVEL • Each sequential step in a food chain or food web, from producer to primary, secondary, or tertiary consumers Energy Flow • Energy flows through an ecosystem in one direction, from sun to producer to consumer to decomposer • Energy used for metabolic purposes at a given trophic level is unavailable to the next trophic level Solar Energy 30% reflected back into space immediately Less than one-billionth of the sun’s total energy reaches Earth’s outer atmosphere. 47% absorbed by the atmosphere Less than 1% drives the winds and ocean currents All solar energy is ultimately reradiated to space as heat 0.02% captured by photosynthesis Fig. 26-14, p. 528 Trophic Levels Energy from sun First Second Third Fourth tropic level trophic level: trophic level: trophic level: Decomposers producers: primary secondary tertiary (saprotrophs) consumers consumers consumers Heat Heat Heat Heat Heat Fig. 26-15, p. 529 Energy from sun First Second Third Fourth tropic level trophic level: trophic level: trophic level: Decomposers producers: primary secondary tertiary (saprotrophs) consumers consumers consumers Heat Heat Heat Heat Heat Stepped Art Fig. 26-15, p. 529 A Food Web Energy Pyramid Decomposers (5060) Tertiary consumers (21) Secondary consumers (383) Primary consumers (3368) Producers (20,810) Fig. 26-17, p. 531 LEARNING OBJECTIVE 9 • Describe the main steps in the carbon, nitrogen, and hydrologic cycles KEY TERMS • BIOGEOCHEMICAL CYCLE • Process by which matter cycles from the living world to the nonliving, physical environment and back again The Carbon Cycle • Carbon dioxide gas (CO2) enters plants, algae, and cyanobacteria • • Which incorporate it into organic molecules through photosynthesis Cellular respiration, combustion, erosion of limestone return CO2 to water and atmosphere • Making it available to producers again Carbon Cycle Air (CO2) 2 Soil 2 2 Animal and microorganism Decomposition plant respiration respiration 1 Photosynthesis by land plants 5 Combustion (human and natural) Marine plankton remains Soil Partly decomposed plant remains 3 Coal 4 Natural gas 6 Dissolved Erosion of CO2 in water limestone to Shells of marine form dissolved organisms: Coal CO2 burial and compaction 7 to form rock (limestone) Oil Fig. 26-18, p. 531 The Nitrogen Cycle 1. Nitrogen fixation • Conversion of nitrogen gas to ammonia 2. Nitrification • Conversion of ammonia or ammonium to nitrate 3. Assimilation • Conversion of nitrates, ammonia, or ammonium to proteins, chlorophyll, and other nitrogencontaining compounds by plants The Nitrogen Cycle 4. Ammonification • Conversion of organic nitrogen to ammonia and ammonium ions 5. Denitrification • Conversion of nitrate to nitrogen gas Nitrogen Cycle Atmospheric nitrogen (N2) 1 Biological nitrogen fixation Nitrogen fixation (nitrogen-fixing bacteria from human activity in root nodules and soil) 5 Denitrification (denitrifying bacteria) 4 Decomposition (ammonification by ammonifying bacteria) Plant and animal proteins Internal cycling (nitrification, assimilation, Assimilation (nitrates, ammonia, or ammonification on land) ammonium absorbed by roots and used to make organic compounds) Ammonia (NH3) and ammonium (NH4+) 2 Nitrification (Nitrifying bacteria) 3 Nitrate (NO3−) Fig. 26-19, p. 532 Nitrogen Fixation Nodules (a) Root nodules of a garden pea (Pisum sativum), which is a legume. Rhizobium bacteria live in these nodules and fix nitrogen, some of which is used by the host plant. Fig. 26-20a, p. 533 Heterocysts (b) Many cyanobacteria fix nitrogen. Shown is Anabaena, a cyanobacterium with specialized cells called heterocysts where nitrogen fixation occurs. Fig. 26-20b, p. 533 The Hydrologic Cycle • Exchange of water between land, ocean, atmosphere, and organisms • • • Water enters atmosphere by evaporation and transpiration Leaves atmosphere as precipitation On land, water filters through the ground or runs off to lakes, rivers, and ocean Hydrologic Cycle Movement of moist air Condensation Atmosphere (cloud formation) 1 Precipitation to ocean 1 Precipitation on land 2 Evaporation from ocean 2 Evaporation from soil, streams, rivers, and lakes 3 Transpiration from vegetation 4 Runoff to ocean 5 Percolation through soil and porous rock Ocean Groundwater Fig. 26-21, p. 534 LEARNING OBJECTIVE 10 • Distinguish between bottom-up and top-down processes Bottom-Up Processes • Availability of resources such as minerals controls the number of producers (lowest trophic level), which in turn controls the number of herbivores, which in turn controls the number of carnivores Top-Down Processes • Regulate ecosystems from the highest trophic level (consumers eating producers) • An increase in number of top predators cascades down the food web through herbivores and producers Bottom-Up and Top-Down Processes Carnivores Carnivores Herbivores Herbivores Producers Producers Nutrients Nutrients (a) Bottom-up processes (b) Top-down processes Fig. 26-22, p. 535 Animation: Exponential Growth CLICK TO PLAY Animation: Carbon Cycle CLICK TO PLAY Animation: Hydrologic Cycle CLICK TO PLAY