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Natural Selection Darwin vs Lamarck • Lamarck - animals pa on acquired traits • Darwin - individuals are selected for survival by combinations of traits • see page 303 Darwin... • More individuals are born than live to reproduce • individuals vary from one another • individuals compete with each other for resources • individuals with the best combination of traits for the given conditions will survive What does natural selection not account for? Neo-Darwinism or Synthetic theory of evolution • Darwin posited that traits are passed on directly to the next generation • Mendel had already determined that traits are passed along as a matter of chance • the combination of Darwinian natural selection and Mendelian genetics is called the synthetic theory of evolution Evidence of evolution • Fossil record • Comparative anatomy (see Figure 7-17 on page 308) • Biogeography • subcellular Is the fossil record a complete record? Comparative anatomy • Homologous features- similar structure in different species • Analogous features - similar function different structure (convergent evolution) • Vestigial organs Biogeography • Evolution occurs only once • Center of origin • Related species have similar patterns of development (figure 17-12, page 313) Subcellular • DNA is universal • the longer the time since divergence the greater the difference in DNA Microevolution • Population = gene pool • Microevolution- small gradual changes within a population over time. Accounts for differences between population of same species Hardy-Weinberg Principle • This never occurs in nature but helps to understand the forces of evolution. • The frequency of genotypes in a population can be determined /described mathematically • Inheritance itself will not cause a change in allele frequencies Inheritance itself will not cause a change in allele frequencies • Allele frequencies will not occur if • • • • • there is random mating there are no mutations there is no migration there is no natural selection UNDER THESE CONDITIONS THERE IS GENETIC EQUILIBRIUM!!!! Mutation • A random, permanent change in DNA • does not determine the direction of evolution (mutations occur randomly, not in response to need or conditions) • Genetic drift - (founder effect) • Genetic bottleneck - Selection • Stabilizing selection - tends to make population more uniform • Directional stabilization - favors one extreme • Disruptive stabilization - favors both extremes • see page 327 Variation • Genetic polymorphism - two or more alleles for a given gene. • What is the heterozygous advantage? Speciation • Species - members freely breed with one another • Reproductive isolation – temporal isolation (different times) – behavioral isolation (courtship patterns) – mechanical isolation Speciation • Allopatric speciation - geographic separation results in new species • sympatric speciation - divergence in same geographic area (insects)