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Characteristic #6 All organisms respond to changes in external environment while maintaining constant internal environment. • Inside is controlled chemically and physically. • External responses: – Skin tans in response to sunlight – Roots change direction moving towards water – Dogs pant in the heat Evolution Change over time Charles Darwin • A naturalist who served aboard research ship called “The Beagle” in 1831 • Observed bird and animal species in Galapagos • Made observations about species characteristics • Published “On the Origin of Species” • Formulated theory of evolution by natural selection • Birds on different islands had different shaped beaks. • Each beak style seemed perfect for island’s food supply. • Darwin speculated that each species of bird had adapted to utilize the food in its environment. • Darwin explained these changes with his theory of evolution by natural selection Darwin’s Observations Evolution Gradual changes in a population or species over time Natural Selection The way these changes occur Patterns of Evolution Artificial selection When humans breed organisms with certain traits to produce offspring with those traits. Examples: dog species, draft horses, garden variety flowers Artificial Selection Dairy cows have been bred by artificial selection for large udders and high milk production. As a result, many cows have udders so large that they cannot walk without swinging their legs out to the side. Patterns of Evolution Natural selection When organisms that have favorable adaptations are selected by nature survive and reproduce. Examples: colors which blend in with habitat, prehensile tail, beaks adapted to local food sources Adaptations Opposable thumb: thumb that allows grasping and holding Evolution by Natural Selection In nature, organisms with the best characteristics for surviving in an environment will live longer and pass those characteristics on to more offspring. Eventually more of the population will have these adaptive traits. In nature, organisms produce too many offspring for environment to support Individuals will have a variety of characteristics Individuals with good traits for survival will live and reproduce more than others, passing on the good traits to their offspring. The population will eventually change in appearance as this trait becomes more common. Two Types of Natural Selection Divergent Evolution When related species become more and more different due to different niches Adaptive Radiation One type of divergent evolution One species in a new environment changes into many new species, each adapted to a particular food source and way of life. Divergent Evolution of Hawaiian Honeycreepers • A small bird migrated to an island • Island had diverse ecosystem (insects, nuts, berries, flowers & nectar, worms, leaves… • Eventually, many species of birds evolved • Each had features adapted for one niche – Different coloration – Different beak shapes – Different anatomical structures Maui Parrotbill `Amakihi Breaks apart wood and then pries out insect larvae. Eats insects and nectar. `Akohekohe Feed primarily on insects and nectar from `ohi`a blossoms Maui Creeper Eats insects among the leaves and branches. The Po`ouli Finds insects, spider and snails under leaves and bark 'I'iwi Honeycreeper Eats the nectar stored in deep-throated flowers. Homologous structures • Structures from related individuals that have become modified for a particular environment. • Usually result from divergent evolution. Second Type of Natural Selection Convergent evolution When two or more unrelated species become more similar due to similar environments. Organisms that have converged due to similar environments Analogous structures • Structures on unrelated species that are similar due to similar functions. • Analogous structures result from convergent evolution. Analogous Structures Fish/shark and Whale • No common ancestry • Similar body shape and streamlining due to common environment. • Structures have become similar due to convergent evolution. Evidence for Evolution and Relatedness of Species Types of evidence that shows if species are related include: – Fossil evidence – Comparative anatomy – Biochemistry – Chromosomes – Embryology – Microevolution What are Fossils? • Fossils - traces of dead organisms • Fossils can be molds, footprints, imprints, bones or entire organisms Determining the Age of a Fossil • Absolute age: carbon dating relies on radioactive elements, which tell the actual age of the fossil. • Relative age: the age of a fossil can be predicted by its location in a rock column Fossil Evidence • • • Fossils can be found in sedimentary rock (mud, sand or clay). Older fossils are found lower in rock Newer fossils are found closer to the surface Sedimentary Rock Geologic column: a tall segment with many layers of sediment Fossils on the bottom of the column are older than those on the top. Oldest Youngest Using Fossils to Understand Relationships • The bone structure in fossils indicates how closely related organisms may be • The sequence of development the horse was determined by the location in rock and the similarities of the fossils 1. Hyracotherium 4. Merychippus 2. Orohippus 5. Pliohippus 3. Mesohippus 6. Modern day Equus Comparative Anatomy • Similar anatomy indicates closer relationships • Homologous structures show closer relationships Crocodile forelimb Whale forelimb Bird wing Vestigial Structures • Structures present in ancestors but no longer needed. • For example: - appendix Snake Whale - tailbones humans (4) gorilla (?) - ear muscles - pelvic girdle Snakes Whales Biochemistry • ALL living organisms contain the same genetic code – the same four DNA nucleotides (AGCT) – The same 20 amino acids • More closely related species have more similar: – Nucleotide sequences in DNA – Amino acid sequences in their proteins All life uses the same genetic code In every living species, the codon UAC in RNA codes for the amino acid tyrosine Cytochrome C • • Cytochrome c is an enzyme in the mitochondria of the body. Humans have 104 amino acids in cytochrome c • • • • Chimpanzees have the identical aa sequence as humans Dogs differ in 13 of the amino acids Snakes differ by 20 amino acids. The more closely related the species, the more amino acids they will have in common. Chromosomes Closely related species have similar gene sequences on chromosomes • Genetically, chimpanzees are 98.5 percent identical to humans. • However, the differences between the species are profound. • Small gene differences make for huge differences in traits Genetic Differences between Chimpanzees and Humans Embroylogy • Embryos from different species show similar features due to shared ancestry • All vertebrates have – Tails – Gill pouch Chick Human Microevolution watching evolution occur Sometimes, evolution occurs on a small scale that we can watch over a period of a lifetime Microevolution In England, the industrial revolution resulted in heavy pollution and soot-covered surroundings. Eventually the trend was reversed and the pollution was reduced. Researchers studied how a species of moth with two forms adapted to its changing environment. In each environment, the form that was best camouflaged lived longer and had more offspring than the other, and eventually became the most common form. Microevolution watching evolution occur The Moth study is one example of microevolution Physiological adaptations can develop rapidly Non-resistant bacterium Antibiotic Resistant bacterium The bacteria in a population vary in their ability to resist antibiotics. When the population is exposed to an antibiotic, only the resistant bacteria survive. The resistant bacteria live and produce more resistant bacteria. Non-resistant bacterium Antibiotic Resistant bacterium Today, penicillin no longer affects as many species of bacteria because some species have evolved physiological adaptations to prevent being killed by penicillin.