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Chapter 52-55 Ecology Characteristics of a Population • A population is a group of individuals of a single species that simultaneously occupy the same general area. Populations may increase in either of these 2 ways • Population Density: # of individuals per unit (ex: # people/square miles) • Population Dispersion: how individuals are arranged or dispersed (even, clumped, random) • Mark and recapture: used to estimate population size . – Individuals are trapped in an area and captured, marked with a tag, recorded, and then released. – After a period of time has elapsed, traps are set again, and individuals are captured and identified. *Why do we care about estimating a population size? • Additions occur through birth, and subtractions occur through death. – Demography studies the vital statistics that affect population size. – A graphic way of representing the data is a survivorship curve. • This is a plot of the number of individuals in a cohort still alive at each age. –A Type I curve shows a low death rate early in life (humans). –The Type II curve shows constant mortality (squirrels). –Type III curve shows a high death rate early in life (oysters). Fig. 52.3 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Factors that effect reproductive potential • Reproducing earlier in life • Generation time (average age of reproduction) • Gestation Period: the time in which a fetus develops (mammals) (elephants – 22 months!) Population Growth • Population Growth Rate: Birth rate – death rate Biotic Potential: the fastest rate at which a population can grow Reproductive Potential: the maximum number of offspring an individual can produce (in lifetime) Who has a bigger reproductive potential: Bacteria or elephants? Exponential Growth • Exponential Growth: Population significantly increases with each generation Occurs with unlimited resources and no competitors/predators R-selection: growth that occurs when birth rates are high Fig. 52.9 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Carrying Capacity • Carrying Capacity: the maximum population an ecosystem can support. • Logistical Growth: in a population, the initial growth is exponential, then the growth slows and eventually stops • K-selection: growth that incorporates the effect of population density on rate of increase Fig. 52.11 Population Growth Models • Exponential model (red) • idealized population in an unlimited environment (Jcurve); r-selected species (r=per capita growth rate) • Logistic model (blue) •carrying capacity (K): maximum population size that a particular environment can support (S-curve); K-selected species • The logistic population growth model and life histories. – This model predicts different growth rates for different populations, relative to carrying capacity. • Resource availability depends on the situation. • The life history traits that natural selection favors may vary with population density and environmental conditions. • In K-selection, organisms live and reproduce around K, and are sensitive to population density. • In r-selection, organisms exhibit high rates of reproduction and occur in variable environments in which population densities fluctuate well below K. Population life history “strategies” • r-selected (opportunistic) • K-selected (equilibrial) • Short maturation & lifespan • Many (small) offspring; usually 1 (early) reproduction; no parental care • High death rate • Long maturation & lifespan • Few (large) offspring; usually several (late) reproductions; extensive parental care • Low death rate Some questions… • Why do all populations eventually stop growing? • What environmental factors stop a population from growing? • The first step to answering these questions is to examine the effects of increased population density. Two Types of Population Regulation • Density Dependent: A more crowded population causes more deaths (ex: starvation, predation, disease) • Density Independent: A certain percent of a population dies regardless of the population density (ex: severe weather, natural disaster) http://www.youtube.com/watch? v=BSVbdaubxxg&NR=1 • The Demographic Transition. – A regional human population can exist in one of 2 configurations. • Zero population growth = high birth rates – high death rates – where? • Zero population growth = low birth rates – low death rates- where? – The movement from the first toward the second state is called the demographic transition. Fig. 52.21 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Fig. 52.22 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Earth’s Carrying Capacity? • Predictions of the human population vary from 7.3 to 10.7 billion people by the year 2050. – Will the earth be overpopulated by this time? • Wide range of estimates for carrying capacity. – What is the carrying capacity of Earth for humans? – This question is difficult to answer. • Estimates are usually based on food, but human agriculture limits assumptions on available amounts. • Ecological footprint. – Humans have multiple constraints besides food. – The concept an of ecological footprint uses the idea of multiple constraints. • For each nation, we can calculate the aggregate land and water area in various ecosystem categories. • Six types of ecologically productive areas are distinguished in calculating the ecological footprint: – – – – – – Land suitable for crops. Pasture. Forest. Ocean. Built-up land. Fossil energy land. Community structure • Community~ an assemblage of populations living close enough together for potential interaction Interactions • Interspecific (interactions between populations of different species within a community): The Niche • Ecological niche~ the sum total of an organism’s use of biotic and abiotic resources in its environment; its “ecological role” √ fundamental~ the set of resources a population is theoretically capable of using under ideal conditions √ realized~ the resources a population actually uses • Thus, 2 species cannot coexist in a community if their niches are identical Ex: Barnacle sp. on the coast of Scotland • Resource partitioning~ sympatric species consume slightly different foods or use other resources in slightly different ways Ex: Anolis lizard sp. perching sites in the Dominican Republic Relationships • Trophic structure / levels~ feeding relationships in an ecosystem • Primary producers~ the trophic level that supports all others; autotrophs • Primary consumers~ herbivores • Secondary and tertiary consumers~ carnivores • Detrivores/detritus~ special consumers that derive nutrition from non-living organic matter • Food chain~ trophic level food pathway Food Chains This is a food chain Grass Insect Bird Fox Where does the initial energy come from? How does each organism obtain their energy? A food chain is a sequence in which energy is transferred from one organism to the next as each organism eats another organism Food Webs • Food webs are interconnected food chain that show many feeding relationships hawk Bird snake mouse seeds insect pine tree - The arrows show the FLOW OF ENERGY Energy Pyramid • Review 10% law Dominant species and keystone species exert strong controls on community structure • Dominant species are those in a community that have the highest abundance or highest biomass (the sum weight of all individuals in a population). – If we remove a dominant species from a community, it can change the entire community structure. (often plants) Keystone Species • Keystone species: a species that has a major impact and role within an ecosystem. Without them the ecosystem will fall apart. – Sea otter (Decrease Sea Otter Increase sea urchinDecrease Kelp- Decrease Fish spawning Most communities are in a state of nonequilibrium owing to disturbances • Disturbances are events like fire, weather, or human activities that can alter communities. – Some are routine. Fig. 53.16 • We usually think that disturbances have a negative impact on communities, but in many cases they are necessary for community development and survival. Fig. 53.18 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Humans are the most widespread agents of disturbance • Human activities cause more disturbance than natural events and usually reduce species diversity in communities. Give examples Succession • Ecological succession~ changes to an ecosystem over time • Primary~ begun in lifeless area; no soil, perhaps volcanic activity or retreating glacier • Secondary~ an existing community has been cleared by some disturbance that leaves the soil intact (fire) Primary Succession In I Am Legend Secondary Succesion Ecosystems • All of the biotic and abiotic factors in an area Trophic Relationships • The autotrophs are the primary producers, and are usually photosynthetic (plants or algae). – They use light energy to synthesize sugars and other organic compounds. • Heterotrophs are at trophic levels above the primary producers and depend on their photosynthetic output. Fig. 54.1 – Herbivores that eat primary producers are called primary consumers. – Carnivores that eat herbivores are called secondary consumers. – Carnivores that eat secondary producers are called tertiary consumers. – Another important group of heterotrophs is the detritivores, or decomposers. • They get energy from detritus, nonliving organic material and play an important role in material cycling. Primary Production • The amount of light energy converted to chemical energy by an ecosystem’s autotrophs in a given time period is called primary production. (very important to an ecosystem) In aquatic ecosystems, light and nutrients limit primary production • Production in Marine ecosystems. – Light is the first variable to control primary production in oceans, since solar radiation can only penetrate to a certain depth (photic zone). Fig. 50.22 –In the open ocean, nitrogen and phosphorous levels are very low in the photic zone, but high in deeper water where light does not penetrate. Fig. 54.5 • Nitrogen is the one nutrient that limits phytoplankton growth in many parts of the ocean. Fig. 54.6 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings • Production in Freshwater Ecosystems. – Solar radiation and temperature are closely linked to primary production in freshwater lakes. Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings – During the 1970s, sewage and fertilizer pollution added nutrients to lakes, which shifted many lakes from having phytoplankton communities to those dominated by diatoms and green algae. • This process is called eutrophication, and has undesirable impacts from a human perspective. Impacts of Eutrophication • Algal blooms block light available to other plants • When algae die, decompose and deplete dissolved oxygen creating dead zones • Controlling pollution may help control eutrophication. –Experiments are being done to study this process. Fig. 54.8 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings Chemical Cycling • • • • • Biogeochemical cycles: the various nutrient circuits, which involve both abiotic and biotic components of an ecosystem Water Carbon Nitrogen Phosphorus Human Impact • Biological magnification: trophic process in which retained substances become more concentrated at higher levels • Greenhouse effect and climate change : warming of planet due to atmospheric accumulation of carbon dioxide • Ozone depletion: effect of chlorofluorocarbons (CFC’s) released into the atmosphere, deplete ozone • Rainforest destruction • Cause: Overpopulation? Conservation Biology • Conservation biology is a goal-oriented science that seeks to counter the biodiversity crisis, the current rapid decrease in Earth’s variety of life. • Extinction is a natural phenomenon that has been occurring since life evolved on earth. – The current rate of extinction is what underlies the biodiversity crisis. – A high rate of species extinction is being caused by humans. Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings What is biodiversity? • Biodiversity: the total of genetically different organisms in an area A lot of biodiversity: Rainforest Levels of Diversity • Species diversity: # of different species in an area • Ecosystem diversity: the variety of habitats, communities within an ecosystem • Genetic Diversity: different genes within a population • Loss of genetic diversity. – If a local population becomes extinct, then the entire population of that species has lost some genetic diversity. • The loss of this diversity is detrimental to the overall adaptive prospects of the species. • Loss of species diversity. – The U.S. Endangered Species Act (ESA) defines an endangered species as one in danger of extinction throughout its range, and a threatened species as those likely to become endangered in the foreseeable future. • Loss of ecosystem diversity. – The local extinction of one species, like a keystone predator, can affect an entire community. – Some ecosystems are being erased from the Earth at an unbelievable pace. • For example, an area the size of the state of West Virginia is lost from tropical forests each year. Biodiversity: Human welfare • 25% of all medical prescriptions • Genetic variability • Aesthetic and ethical reasons • Species survival Biodiversity crisis- 4 major threats Major Threats: 1. Habitat destruction ~ single greatest threat; cause of 73% of species designation as extinct, endangered, vulnerable, rare; 93% of coral reefs Biodiversity crisis- 4 major threats Major Threats: 2.Competition by exotic (non-native) or invasive species – species that humans move from native locations to new geographic locations - What are the problems associated with this? Biodiversity crisis- 4 major threats Major Threats: 3. Overexploitation ~ commercial harvest or sport fishing; illegal trade, hunting. • The African elephant has been overhunted and the populations have declined dramatically. • The bluefin tuna is another example of an overharvested species. Fig. 55.9 Copyright © 2002 Pearson Education, Inc., publishing as Benjamin Cummings • Disruption of food chains. – The extinction of one species can doom its predators, but only if the predator feeds exclusively on this prey. Conservation biology focus • Preservationism: setting side select areas as natural and underdeveloped Nature Reserves Population & species level conservation • Biodiversity hot spot: small area with an exceptional concentration of species • Endemic species: species found nowhere else • Endangered species: organism “in danger of extinction” • Threatened species: likely to become endangered in the foreseeable future • Bioremediation: use of living organisms to detoxify polluted systems