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Unit 10: Evolution Ch. 12 Evolution & Ch. 13 Adaptation & Speciation 300 History of Earth • How old is Earth? – ~4.6 billion years old • What was early Earth like? – hostile... a lot of volcanic activity & lightning, many meteorite impacts, high levels of UV light • What gases were in the primitive atmosphere? – methane, ammonia, hydrogen, water vapor • What gas was missing? – oxygen Origin of Life • When did life 1st appear? – ~3.5 billion years ago • What did Miller & Urey do in their experiment? – simulated primitive atmosphere & zapped it with electricity • What happened? – ~1 week amino acids & other simple organic molecules formed (in water) » precursors to 1st cells The First Organisms: Heterotrophs • What does the “Heterotroph Hypothesis” or “Primordial Soup Hypothesis” state? – simple compounds formed 1st complex compounds primitive pre-cells prokaryotes (1st cells were heterotrophs) • Why heterotrophic? – b/c no O2 for cellular respiration • What did they feed on? – naturally organic material (primordial soup) in ocean (ex. amino acids) » Similar to some present-day bacteria, they obtained energy from simple chemical reactions. Later Organisms: Autotrophs • What type of organisms followed the prokaryotic heterotrophs? – prokaryotic autotrophs stromatolites • What did they release into the atmosphere? – oxygen (….causing the oxygen revolution!) • Why was oxygen so important? – allowed development of eukaryotes & complex organisms What is Evolution? • theory that species/populations change over time – Can an individual’s genes change? • No… – Can a population’s gene pool (total genes) change? • Yes… – when allele frequencies (%) change due to natural selection 4 Mechanisms (Causes) of Evolution • 1.How does migration cause evolution? – gene frequencies change as individuals move in or out of the population • 2. How does genetic drift cause evolution? – pure CHANCE causes a change in gene pool of a small population 4 Mechanisms of Evolution • 3. How does mutation cause evolution? – random changes in DNA sequence can change amino acid sequence & protein coded for… & can change how the trait is expressed • Do all mutations matter in evolution? – No… to matter they must be in gametes » …able to be passed down (heritable) 4 Mechanisms of Evolution • 4. How does natural selection cause evolution? – organisms with the variation best suited for environment (adaptation) tend to survive, & reproduce more (“survival of the fittest”) • What will that cause? – favorable adaptation will be passed to more offspring than before – acts on individuals, BUT causes evolution of population 4 Mechanisms of Evolution •Which trait was a better adaptation and selected for? •How were frequencies of different alleles affected? Explanations for How Evolution Happens • What did Jean Baptist Lamarck propose about how evolution occurred? – “Theory of Acquired Characteristics” (use vs. disuse) • acquired traits are passed on & leads to changes in species – Proposed that nature drove organisms from simple to increasingly complex forms • Did experimentation support Lamarck’s thinking? –no Explanations for How Evolution Happens • Charles Darwin (naturalist on HMS Beagle) – What did he do on his trip? • collected fossils & specimens – observed tremendous biodiversity » noticed unique organisms similar to organisms in other parts of world… determined they were all related Climate varied from island to island. intermediate shell domed Shell large rainfall saddlebacked shell desert Explanations for Evolution • What did Darwin conclude from his observations? – life forms can & do change…. but, questioned how this change occurred… •“answers” pointed to evolution or “descent with modification” –needed to test ideas before explaining how or why such changes happen Explanations for Evolution • When thinking about HOW evolution happened, Darwin noted artificial selection (selective breeding)… – plan/goal = to produce preferred offspring • Darwin wondered, “Is there a force in nature that simulates artificial selection???” Darwin’s Explanation: Natural Selection • What were the 4 main points Darwin used to explain how evolution occurred? 1859 1. Overproduction of offspring 2. Struggle to survive (competition for existence) 3. Variation within population 4. Successful reproduction (“survival of the fittest”/ best adapted organisms are “naturally selected”) Video: Who Was Charles Darwin? Darwin’s Explanation: Natural Selection 1. Overproduction of offspring – Why is this necessary? • to ensure that at least some offspring will survive long enough to reproduce & pass on genes Darwin’s Explanation: Natural Selection 2. Over production leads to the “struggle to survive”… How? – more organisms than resources to go around leads to competition (idea based on work of Thomas Malthus) • Organisms compete for…? – food, shelter, mates, space, light, water, nutrients, etc. • Population levels off at the “carrying capacity”. – maximum # that can be supported given available resources Video: Darwin & Malthus Darwin’s Explanation: Natural Selection •The survivors of the “struggle to survive”… camouflage – Who are they? • individuals w/ variations that are best suited for that particular environment (adaptations) – How do they get these traits? • chance & variation – unlike artificial selection which has plan/goal – Were these traits a response to their environment? • NO… it just so happens that the variation is beneficial – & allows for increased survival & reproduction of more offspring with that trait (due to natural selection for that trait) Darwin’s Explanation: Natural Selection Darwin’s Explanation: Natural Selection Contractile Vacuole Video Darwin’s Explanation: Natural Selection 3. In any given population, there will be a variation within a trait. Darwin’s Explanation: Natural Selection •What causes variations within a population? – Sexual reproduction – Crossing over during meiosis – Arrangement of chromosomes during meiosis (independent assortment) – Mutations • Sexual repro. & meiosis speed up evolution… increases variety • Darwin did NOT have knowledge of genetics! Darwin’s Explanation: Natural Selection 4. How does natural selection lead to successful reproduction? – Those organisms best suited for environment tend to survive, reproduce, & pass on genes (“survival of fittest”) • more offspring have favorable adaptation than before Darwin’s Explanation: Natural Selection • Ex. Pepper Moths – England – Original color – 99% white (trees w/ light bark) • Industrial Revolution, 1850s – produced soot – Enviro. changes - 1890s - 99% black (trees w/ dark bark) Pepper Moth “Game” http://www.techapps.net/interactives/pepperMoths.swf Principles of Natural Selection "Interactive" Darwin’s Explanation: Natural Selection Adaptive Radiation Hawaiian Honeycreepers How would Darwin adjust the description given by Lamarck about the length of giraffes necks? Lamarck vs. Darwin Darwin’s Explanation: Wallace’s Same Conclusion • Who was Alfred Wallace (1858)? – naturalist that arrived at same basic explanation as Darwin • However, Darwin usually given credit – but BOTH men should be credited Indirect Evidence to Support the Theory of Evolution: Fossil Record • fossil record – evidence of: • change in organisms over time Which layer is youngest? oldest? Explain… Law of Superposition • age of the Earth Indirect Evidence to Support the Theory of Evolution: Fossil Record • Why is the fossil record limited? – decomposition – type of material • What fossilizes best? – hard parts…. teeth, bones, scales – incomplete record • Why? – – not all organisms are fossilized not all fossils are found – easily disrupted • Why? – some are destroyed by natural processes or people Indirect Evidence to Support the Theory of Evolution: Comparative Anatomy • comparative anatomy – compares structures of different organisms living today 1. homologous structures 2. analogous structures 3. vestigial structures Indirect Evidence to Support the Theory of Evolution: Comparative Anatomy • 1. What are “homologous structures”? – similar structures with modifications (ex. location & arrangement of bones) • Which are evidence of …? – – a common ancestor later organisms becoming more different (divergent evolution) Indirect Evidence to Support the Theory of Evolution: Comparative Anatomy • 2. What are “analogous structures”? – different structure, but same function (ex. wings made of different materials… bird = bone vs. insect = chitin) • Which are due to…? – similar environment/use » but… do NOT have common ancestor » later organisms become more similar (convergent evolution) Indirect Evidence to Support the Theory of Evolution: Comparative Anatomy • 3. What are “vestigial structures”? – structures with reduced size or function that often have no apparent use • Which are evidence that…? – species has changed Indirect Evidence to Support the Theory of Evolution: Comparative Embryology • How does embryonic development provide evidence for evolution? – early stages very similar • later stages more distinction – What does this tell us? » organisms inherited same basic body plan from common ancestor » greater # similar stages more closely related Indirect Evidence to Support the Theory of Evolution: Comparative Biochemistry • What codes for similar organic compounds (ex. amino acids & proteins)? – similar DNA • DNA most reliable form of evidence – What is the significance of 2 very different species sharing large amounts of the same DNA? • evidence of common ancestry – higher %, more closely related Direct Evidence to Support the Theory of Evolution: Observation • Ex. of rapid evolution that allow us to observe evolution…? 1. antibiotic- resistant bacteria • 2. • MRSA pesticide-resistant insects Resistance already present in some individuals (variation) – resulting from mutation Evolution & Classification • Evolutionary relationships are the basis of classification – closely related share more groups (similar classification) • black bear & brown bear – same down to genus – distantly related share fewer groups (dissimilar classification) • snake & brown bear – same only to phylum DOMAIN Eukaryota