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EVOLUTION Evolution - change over generations Organisms change over time – this is a fact. How organisms have changed throughout earth’s history is theory. EVIDENCE OF EVOLUTION Fossils Comparative anatomy Embryology Biochemistry Genetics Direct observation FOSSILS Fossils - a trace of an organism that lived long ago Fossil records- use radiometric dating or rock layers to date fossils Fossil records are incomplete: • soft tissues • weather Geographic distribution of species Similar organisms in different areas Fossils of extinct organisms COMPARATIVE ANATOMY Comparative anatomy- The study of structures of different organisms Homologous structures - similar structures modified between different organisms • The more similar the structures, the closer the relationship between two organisms • Example: a bat’s wing, a horse’s leg and a human forearm Homologous structures Vestigial Organs - structures that have no function in present organisms, but may have been used in ancestors Example in animals • pelvic bone in some snakes and whales Examples of vestigial organs in humans • appendix • wisdom teeth • body hair • pointed canines A transitional fossil EMBRYOLOGY Comparative embryology - the study of embryos (early stages of life) of different organisms The more similar the embryos, the more closely related two organism are • ex. human and pig embryos appear very similar to each other BIOCHEMISTRY & GENETICS Chemicals (like proteins, DNA, mRNA etc) in different organisms show similarities. More similar chemicals means more closely related Example- Hemoglobin in chimpanzees is very similar to hemoglobin in humans GENETICS How are offspring genetically different from their parents? In other words, how can evolution occur from one generation to the next? Meiosis – remember, individuals can make lots of genetically different sex cells Mutations – DNA can change! Sex – combines two individuals’ DNA together DNA sequences of all living things have many similarities Base sequences of DNA • DNA of humans and chimpanzees is 98% identical • DNA of humans and other mammals is 80% identical Hemoglobin sequences in vertebrates DIRECT OBSERVATION Direct observation - evolutionary changes are happening NOW • Example - bacteria becoming resistant to antibiotics Charles Darwin in 1859, the year The Origin of Species was published CHARLES DARWIN Charles Darwin – an Englishman credited for first describing natural selection • HMS Beagle – a British survey ship on which Darwin was a naturalist. This ship sailed around the world in 1831. Figure 15–1 Darwin’s Voyage Section 15-1 Galapagos Islands – located 600 miles west of South America. The islands have very different climates and thus the organisms adapt according to the environment. Giant tortoise, marine iguanas, and finches vary from island to island due to different environments Giant Tortoises of the Galápagos Islands Pinta Pinta Island Tower Marchena Intermediate shell Fernandina James Santa Cruz Isabela Santa Fe Hood Island Floreana Isabela Island Dome-shaped shell Hood Saddle-backed shell A Galápagos Islands tortoise Artificial Selection – people chose particular species to breed (selective breeding) Artificial selection helped to explain that variations in species could make them better adapted. Dog breeders, farmers, gardeners have been using artificial selection for years. Golden Doodle Cockapoo Himilayan Cocker spaniel and poodle Golden retriever and poodle Persian and Siamese Tangelo = madarin orange and grapefruit Broccoliflower = broccoli and cauliflower Artificial selection: diverse vegetables derived from wild mustard NATURAL SELECTION: The survival of the fittest Darwin wondered why there were so many different species of finches on the Galapagos Islands. He proposed the concept of natural selection. Natural Selection - the best adapted individuals in a population survive and reproduce offspring that are also well adapted The least adapted produce fewer offspring and are less likely to survive. Among the various traits that exist, certain ones may benefit a population The environment typically does not cause the variation! • Examples • Virus from outer space – kills blueeyed people (but does not change a person’s eyes to blue) • Spadefoot toad – must bury itself in the ground and mate quickly when it comes to the surface, therefore, it has a loud croak and long toes! TYPES OF SELECTION Directional Selection – when individuals at one end of the curve have higher fitness than individuals in the middle. Example: birds with larger beaks are better able to survive food shortages than those with small and medium beaks Figure 16–6: Graph of Directional Selection Key Directional Selection Low mortality, high fitness Food becomes scarce. High mortality, low fitness Stabilizing Selection - when individuals at middle of the curve have higher fitness than individuals at the ends. Example – average-sized babies are most likely to survive compared to really big and really small babies Figure 16–7: Graph of Stabilizing Selection Stabilizing Selection Key Low mortality, high fitness High mortality, low fitness Birth Weight Selection against both extremes keep curve narrow and in same place. Disruptive selection - when individuals at both ends of the curve have higher fitness than individuals in the middle. Example: due to small and large seeds, birds with small and large beaks are most fit compared to birds with medium beaks Figure 16–8: Graph of Disruptive Selection Disruptive Selection Low mortality, high fitness High mortality, low fitness Population splits into two subgroups specializing in different seeds. Beak Size Number of Birds in Population Key Number of Birds in Population Largest and smallest seeds become more common. Beak Size Disruptive selection in a finch population Types of selection Some vocab… Gene Pool - the entire collection of genes among a population Population - All organisms within a breeding group (same species) Gene Frequency - The number of times an allele appears in a gene pool Figure 16–2: Relative Frequencies of Alleles Sample Population 48% heterozygous black 16% homozygous black 36% homozygous brown Frequency of Alleles allele for brown fur allele for black fur Changes in gene frequency mean that the population has evolved. An entire population evolves, not an individual! A specific gene may be “selected” for by the environment. GENETIC DRIFT Genetic Drift - rapid changes in gene frequency of a small, isolated population Example: in small populations, a recessive allele can become much more common In the Amish population, certain genetic disorders are more common than in the general population. Genetic Drift Sample of Original Population Founding Population A Founding Population B Descendants Genetic Drift Sample of Original Population Descendants Founding Population A Founding Population B OCEANIC ANGLER FISH Sexual dimorphism The deep sea angler shows a very strange sexual dimorphism. The male is very small and attaches itself to the body of the female. The teeth and the jaw recedes and the blood circulating of the two animals become one. The male spends the rest of his life attached to the female. ADAPTATIONS Adaptations - physical traits of a living organism that help it to survive in its environment Adaptations evolve by chance! TYPES OF ADAPTATIONS Structural – physical parts • Ex. woodpecker’s tongue, anglerfish, anteater’s snout, hummingbird’s beak Physiological – chemicals made by organism • Ex. poison venom of a snake, ink of a squid Behavioral Ex. migration of birds, mating dances Buffalo Bill Dam on the Shoshoni River at Cody, Wyoming and those dots are bighorns WALKING ACROSS IT. TYPES OF EVOLUTION Convergent Evolution – when species that are not closely related evolve similar traits (two species look like they are closely related and they are not) • Example: dolphins (mammals) and sharks (fish) Divergent Evolution – one species evolves into two or more species with different characteristics Example: lions and tigers evolved from a common ancestor Liger Speciation Speciation –evolution of two different species from one common ancestor. What keeps two species apart so that they evolve differently? Isolating Mechanisms – keep groups apart Geographic Isolation – geologic changes occur that isolate individuals of a population – Ex. Squirrels separated by Grand Canyon and Darwin’s finches separated by being on different islands Squirrels in the Grand Canyon Figure 15–14: Geographic Distribution of Living Species Beaver NORTH AMERICA Muskrat Muskrat Beaver and Muskrat Coypu Capybara Capybara SOUTH AMERICA Coypu Coypu and Capybara Genetic Isolation – genes don’t mix right – Donkeys and horses make mules, but mules are sterile. Temporal isolation – timings off – Different species of skunks that mate at different times of the year. Behavioral isolation – behaviors don’t match – Certain species of lightning bugs use different blinking patterns to attract mates. A male bower bird builds a “bower” to attract mates. Blue footed boobies doing their mating dance Birds of Paradise doing their mating dance Many different mating scenarios THEORY OF ORIGIN OF LIFE Theory - a major hypothesis that has withstood the test of time • Based on observations and evidence • The closest to a complete explanation that science can offer • 40 second evolution • Homer Simpson Evolution ORIGINAL ATMOSPHERE Hydrogen cyanide, carbon dioxide, carbon monoxide, nitrogen, hydrogen sulfide, and water No oxygen gas FIRST ORGANISMS Prokaryotes [bacteria] - first cells; have genetic material (RNA or DNA); starting 3.5 bya First prokaryotes were heterotrophs (eat food) Later, some became autotrophs (make own food) Chemosynthesis - using energy from chemicals to make food Photosynthesis - using CO2 and light to make food • Cyanobacteria put oxygen into atmosphere. • Oxygen drove some life forms to extinction, while other life forms evolved new ways to use oxygen (aerobic respiration). • Oxygen formed ozone layershielded earth from harmful UV and allowed an explosion of life (2.7 bya) A painting of early Earth showing volcanic activity and photosynthetic prokaryotes in dense mats Other big steps… Eukaryotes (cells with nuclei) about 2.2 bya Plants and animals begin in water Multicellular organisms and sexual reproduction in plants and animals (aquatic life) Increased genetic diversity Plants and later animals colonize land around 4.5 mya. First dinosaurs and then first mammals (245 mya) Dinosaurs go extinct around 65 mya and this allows for evolution of larger mammals. Extinctions are often followed by explosions of new life forms! Remember your eras: Precambrium. Paleozoic, Mesozoic, and Cenozoic. HUMAN EVOLUTION Did man come from Monkey? NO! Humans and monkeys evolved from a common ancestor that lived approximately 65 million years ago and was a tree dweller (roughly the size of a cat). This was not a monkey like the monkeys today. Figure 32–16: Human and Gorilla Skeletons Comparing Human and Gorilla Skeletons Modern Human Modern Gorilla Modern Human Modern Gorilla Skull atop S-shaped spine Skull atop C-shaped spine Spinal cord exits at bottom of skull Spinal cord exits near back of skull Arms shorter than legs; hands do not touch ground during walking Arms longer than legs; hands touch ground during walking Pelvis is bowlshaped Pelvis is long and narrow Thigh bones angled inward, directly below body Thigh bones angled away from pelvis Evolution of Primates Anthropoids developed into 3 groups: New World Monkeys – includes squirrel and spider monkeys; have prehensile tail (can coil around branches) and lived in trees; found in central and S. America Old World Monkeys – include giant apes, baboons, and macaques; do not have prehensile tail, but spend a lot of time in trees; found in Africa and Asia Hominoids (great apes) – include orangutans, gorillas, chimpanzees, and humans HOMINIDS Hominid family includes modern humans and several distinctly different extinct species Bipedal – two-foot locomotion Freed hands to use tools Opposable thumb – allowed hand to grasp objects and tools Increased brain size Family that walks on all four Comparison of Skulls of Human Ancestors Large brow ridge Large nose Large canine teeth Face protrudes forward Australopithecus afarensis Weak brow ridge Homo erectus Large brain case Round, high skull Inflated cheeks Large nose Even teeth Strong chin Neanderthal Cro-Magnon Human Evolution Shockwave video Modern Homo sapiens Australopithecus afarensis Very small brain capacity ~3.9-3.0 million years ago (mya) “Lucy” Bipedal apes that spent some time in trees Short (Lucy is 1m) Homo habilis “Handy man” ~2.5 mya Increased brain size Began to make and use tools 1.3 meters tall (4 feet tall) Homo erectus “Upright man” ~ 1.8 mya - 250,000 years ago Taller Larger brain capacity (comparable to modern humans) Built fires Migrated out of Africa into China and Asia Homo sapiens (“wise man”) Homo sapiens neanderthalis Neanderthals Overlapped in existence with modern humans Heavier brow ridge Slightly larger brain capacity than modern human Lived ~200,000 – 30,000 years ago Homo sapiens sapiens First arose in Africa around 100,000 years ago. Lived with Neanderthals for ~50,000 years. Tools, art, and buried dead with rituals One group became Cro-Magnon (in Europe) ~40,000 years ago. We are Homo sapiens sapiens