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2. Early history of Earth 1. 4.6 billion years old 1ST life 3.5 billion years ago Fossils – any evidence of an organism that lived long ago Classified by the way they are found Casts Trace Fossils Imprints Mold Petrified Fossils Preserved (amber) or frozen (whole organisms) Fossil Pollen CASTS TRACE FOSSILS Markings or evidence of animal activities (ex. Footprints) Mold of organism is created & becomes filled in by minerals in the surrounding rock Produces a replica of the original organism IMPRINTS MOLD Fossils form before sediments harden into rock Organism is buried – it decays & leaves an empty space in the rock that is the exact shape of the organism PETRIFIED FOSSILS Hard parts of organisms are sometimes penetrated and replaced by minerals, atom-foratom When minerals harden, an exact stone copy of the original organism is produced PRESERVED OR FROZEN Entire intact organism can be found frozen in ice or preserved in fossilized tree sap Rare but valuable! because even the most delicate parts of the organism are usually preserved Fossil Pollen gives clues to ancient climate and environmental conditions Fossils are found in sedimentary rocks (small pieces of sand/mud/clay Relative Dating – sedimentary layers closer to Earth’s surface are “younger” Organisms are related to the layer they’re found Radiometric Dating – finds absolute age of fossils Uses isotopes that break down over time 1ST of January – Earth formed (4.6 bya) March 20 – Precambrian Era begins - oldest evidence of life Mid-October (!!) – end of PreCambrian Era Algae, sponges, jellyfish Mid-October – Paleozoic Era begins EXPLOSION of life Land plants! Earliest vertebrates amphibians & reptiles Continental drift separates continents Triassic = 1ST small mammals Jurassic = (mid-December) – Age of Dinosaurs Cretaceous = larger mammals & flowering plants Current era “December 31” – Humans Appear Fossil record shows evidence of many extinctions Early Ideas Spontaneous generation – life was thought to be produced from nonliving matter Experiments of Francesco Redi (1668) Disproved this theory for large organisms Louis Pasteur (1800’s) disproved the belief that microorganisms grew spontaneously from the air Biogenesis = important law of biology living organisms come only from other living organisms Question to be answered: How did life begin on Earth? Answers are based on experimental evidence… not fossils Alexander Oparin (1930’s) – proposed theory that life began in the oceans (“primordial soup”) Stanley Miller & Harold Urey (1953) – simulated early Earth’s atmosphere; created amino acids and organic compounds Small pools of warm water theorized to be sites for production of proteins, ATP, and nucleic acids… which become surrounded by a membrane Sidney Fox – produced protocells (large ordered structure that carried out some activities of life such as growth, division, & metabolism) 1ST cells anaerobic heterotroph Prokaryotes – simple cells that did not use Oxygen and got their food from their surroundings Over time chemosynthetic autotrophs appeared… could make their own food! Similar to present day Archaebacteria that live in harsh conditions (will learn about those later) Finally – photosynthetic prokaryotes; 3.5 byo fossils have been found in Australia These cells produced Oxygen & started to change Earth’s atmosphere; leading to evolution of aerobic respiration “oxygen revolution” Presence of oxygen led to development of Ozone (O3) layer that shielded Earth from UV rays & allowed for evolution of more complex organisms Charles Darwin & Natural Selection Development of evolutionary theory – 1. geologic evidence showed Earth is very old 2. Fossil evidence pointed to changes in life forms over time 3. Scientists wondered where species came from, and how they changed over time 4. Charles Darwin proposed an explanation that has become accepted by the majority of scientists. Founder of modern evolutionary theory 1831- naturalist aboard H.M.S. Beagle spent 5 years on expedition to South America & South Pacific Ocean Visited Galapagos Islands volcanic islands 1000km off western coast of Ecuador, SA Observed & gathered many specimens Back in England – influenced with population studies & idea of struggle for survival Conducted his own experiments with artificial selection = breeding for specific traits 1859 – proposed mechanism of natural selection for evolution Stabilizing Selection 1. Favors average individuals 2. Ex. Spider size = too large can be captured/eaten! Too small can’t hunt well Directional Selection 1. Favors one of the extreme forms of a trait 2. Ex. Woodpeckers = looooong beaks may be more favorable if insects live deep within trees Disruptive Selection 1. Favors both extreme forms of a trait 2. Ex. Limpet (mollusk) = both white & black forms can hide on rocks; grey-colored forms are visible & tend to get eaten by birds Geographic Isolation Physical barrier separates a pop. into groups This is what happens on islands Reproductive Isolation Organisms that once interbred are prevented from producing fertile offspring May be the result of seasonal differences in mating Polyploid Speciation Results from chromosomes not separating during meiosis Results in an immediate new species common in plants Gradualism Adaptive changes over time Supported by the fossil record Example camels & horses Punctuated Equilibrium Rapid bursts of speciation with stable periods in between May result from climate change Also supported by the fossil record Example elephants Adaptive Radiation Process of evolution of an ancestral species into many species that occupy different niches Niche = role a species plays in a community regarding food, space, reproduction, and how it interacts w/nonliving factors EX. Honeycreepers (birds) of Hawaiian Islands Similar in body size & shape VERY different in plumage & beak shape Species begin to adapt to different environmental conditions Species become less & less alike The common ancestral mammal probably existed about 190 m.y.a & with the extinction of the dinosaurs, underwent extensive adaptive radiation to fill a wide range of ecological niches Distantly related organisms evolve similar traits Response to similar environments & selection pressures Dolphins & fishes (yes this is correct if you’re talking about different varieties/species) Population Genetics & Evolution 1. within the lifetime of one individual, new features cannot evolve 2. Natural selection operates only on populations over many generations B. Vocabulary for Population Genetics 1. Gene Pool = entire collection of genes in a population 2. Genetic Equilibrium = population in which the frequency of alleles does not change from generation to generation 3. Frequency = how often a particular allele occurs in the gene pool Mutation – rare that mutations result in favorable variation Genetic Drift Changes in allelic frequency by chance processes More likely to occur in small populations why? In small populations, individuals with recessive genes are more likely to mate Ex. High incidence of polydactyly in an Amish population in Pennsylvania Migration of individuals in & out of populations Variations exist within populations Some variations are more advantageous for survival & reproduction that others Organisms produce more offspring than can survive Over time, offspring of survivors will make up a larger proportion of the population Depending on environmental factors, after MANY generations, a population may come to look very different from the original population Structural adaptations arise over MANY generations Examples of Structural Adaptations 1. Mimicry = an organism copies, or mimics, another Monarch 2. Camouflage = an organism can blend in with it’s butterflies surroundings have a bitter awfultaste 3. to Physiological Adaptations CAN YOU SEE THE birds… - changes in metabolic___________________? processes Viceroy’s don’t produce the - occurs very quickly in bacteria same chemical, Antibiotic resistance but birds learn not to eat them based on the pattern Fossils Anatomical Studies 1. Homologous Structure Modified structure that is seen among different groups of descendents EX. Forelimbs of crocodiles, bats & chickens Similarity of structure means organisms are closely related 2. Analogous Structure Similar in function, but different in structure EX: Birds & butterfly wings have the same job (flight) but are made of different materials 3. Vestigial Structure Structure without a function; shows structural change of a species over time EX. Appendix; flightless birds; eyes of sightless cave fish Studies of Embryos similarities in embryo appearance Which one is the human? Fish? Bird? Rabbit? Genetic Studies similarities in DNA patterns suggest close ancestors