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C4 and CAM Photosynthesis Adaptations to high temperature C4 Photosynthesis • Some plants, adapted to hot, dry environments, use a special process called C4 photosynthesis. • Some examples of C4 plants are: Corn, Sugar Cane, CrabgrassAND Most tropical grasses are C4 plants. C3 Plants Plants that use only the Calvin cycle for light independent reactions may be referred to as C-3 because during the cycle, the 6-C sugar splits to two 3-carbon sugars. Specialized leaves of a C-4 plant have tightly packed layers of cells. They have bundle sheath cells and mesophyll cells surrounding leaf veins. What occurs during C-4 photosynthesis? Carbon Dioxide is first incorporated into a C-4 (four carbon) acid in the mesophyll cells. Why is C-4 necessary? When stomata close… When this happens photosynthesis operates at a much reduced rate, due to RuBP bonding with Oxygen under low CO2 conditions. Thus some of the enzyme is converted to CO2, bypassing the Calvin cycle. When this happens photosynthesis operates at a much reduced rate. The rate of photorespiration increases C4 photosynthesis, therefore provides the needed carbon dioxide when the stomata close. It is a specialization discovered in desert plants such as cactus, and pineapple. Other C4 plants are: Jade plant and agave plant. • C4 plants can function at maximum efficiency in conditions of high temperature and high light intensity while keeping their stomata closed to avoid excessive water loss. • CAM evolved in plants that grow in extreme heat & little rain. (It is not very efficient, and plants grow slowly.) • By contrast, high temperature may inhibit photosynthesis in C3 plants by as much as 40%! What Happens in CAM Photosynthesis?