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Chapter 5 Key Concepts • To understand living organisms, one must have a basic understanding of the variety of compounds from which organisms are built. • Four groups of macromolecules are necessary for life: carbohydrates, lipids, proteins, and nucleic acids. • All living organisms are composed of cells. • Cells can be either prokaryotic or eukaryotic • Cells produce new cells by the process of cell division • Evolution is the process by which the genetic composition of populations of organisms changes over time • Natural selection favors the survival and reproduction of those organisms that possess variations that are best suited to their environment • A species is a group of physically similar, potentially interbreeding organisms that share a gene pool, are reproductively isolated from other such groups, and are able to produce viable offspring. • The binomial system of nomenclature uses two words, the genus and the species epithet, to identify an organism. • Most biologists classify organisms into one of three domains, categories that reflect theories about evolutionary relationships. • Phylogenetic trees and cladograms indicate evolutionary relationships among groups of organisms Building Blocks of Life • Macromolecules – carbohydrates – lipids – proteins – nucleic acids Carbohydrates • Contain C, H and O • Sugars and polysaccharides most common carbohydrates • Sugars – monosaccharides • ribose and deoxyribose are in nucleic acids • glucose is the basic fuel molecule for cells – disaccharides • sucrose • maltose • lactose • Polysaccharides – polymers – storage forms • starches, made of glucose • glycogen, “animal starch” – structural polysaccharides • cellulose • chitin Lipids • Fats, Oils & Waxes • Composed of C and H – fatty acids – Triglycerides • Functions within marine organisms – store energy, cushion organs, buoyancy – cell membranes – steroids – waxes Proteins • Proteins are polymers of amino acids – 20 amino acids – polypeptides • Functions of proteins – muscles and connective tissue – enzymes—biological catalysts – transport or store chemicals Nucleic Acids • DNA (deoxyribonucleic acid) – Large, double stranded, helix-shaped molecule • N-containing base = adenine, guanine, cytosine or thymine • contains genes (genetic material) • genes direct synthesis of proteins • can copy itself so that genes can be past from one generation to the next • RNA (ribonucleic acid) – usually a single-stranded molecule • sugar = ribose • N-containing base = adenine, guanine, cytosine or uracil – functions in protein synthesis • messenger RNA (mRNA) • ribosomal RNA (rRNA) • transfer RNA (tRNA) Cells • All cells are capable of: – metabolism – growth – Reproduction • • Cell membrane Cytoplasm, within the cell membrane is composed of cytosol and organelles Organelles • Nucleus and ribosomes – Nucleus: contains the cell’s DNA and acts as the control center – Ribosomes – endoplasmic reticulum • rough ER • smooth ER – Golgi apparatus – lysosomes – Vacuoles – chloroplasts – mitochondria – Flagella – Cilia Energy Transfer in Cells • Photosynthesis – low-energy molecules combine to form high-energy food molecules – chloroplasts • Cellular respiration – releases energy from food molecules – most occurs within mitochondria – food molecules are broken down to create ATP and release CO2 as a waste product Cellular Reproduction • Cell division in prokaryotes – single chromosome – binary fission • Cell division in eukaryotes – Mitosis – Cytokinesis Levels of Organization • cells are grouped into tissues • different tissues combine into organs • groups of organs make up organ systems Evolution and Natural Selection • Evolution—the process by which populations of organisms change over time • Evolutionary biology investigates: – how and when organisms evolved – what role the environment plays in determining the characteristics of organisms that can live in a given area Darwin and the Theory for Evolution • Voyage of discovery – Darwin traveled on the HMS Beagle for 5 years, beginning in 1831 – Darwin was influenced by Charles Lyell and other geologists who concluded that: • since geological change is slow and continuous, the earth is very old • slow and subtle changes become substantial when they continue for centuries/millennia • Formulating a theory for evolution – Darwin was inspired by Thomas Malthus – Darwin developed his hypothesis “evolution by natural selection” – Published On the Origin of Species by Means of Natural Selection • Theory of evolution by natural selection – artificial selection – similar process was occurring in nature? – natural selection favors survival and reproduction of those organisms best suited to their environment Four basic premises of Darwin’s theory • All organisms produce more offspring than can possibly survive to reproduce. • Variation in inherited traits • Resources are limited • Traits that make individuals better adapted to their environment are more successful, and pass on those traits to their offspring. Genes and Natural Selection • Modern evolutionary theory – Darwin’s 1858 idea refined by modern genetics – genes • produce traits • can exist in alleles • the offspring receives 1 allele for a trait from each parent, producing many possible combinations • Role of reproduction – in asexual reproduction, offspring are clones – variation results from mutation only – in sexual reproduction, chromosomes from 2 parents are combined – meiosis forms haploid cells called gametes • Population genetics – Organisms adapt to survive – ability to adapt is limited by the gene pool – Only individuals that have combinations of genes and alleles that allow adaptations to their surroundings are likely to survive and reproduce – Reproduction is success Evolution of New Species • Typological definition of species – based on morphology • Modern species definition – a species is one or more populations of potentially interbreeding organisms that are reproductively isolated from other such groups – reproductive isolation: members of a different species are not in the same place at the same time or are physically incapable of breeding, so genes from different species are not mixed – isolating mechanisms that prevent fertilization • habitat isolation • anatomical isolation • behavioral isolation • temporal isolation • biochemical isolation Post fertilization isolating mechanisms • incompatible genes or biochemical differences • the hybrid offspring may survive but be infertile or weak Allopatric speciation Classification: Bringing Order to Diversity • Linnaeus and the binomial system • Taxonomic categories – Early schemes of classification • all living things were classified into 1 of 2 kingdoms, Animalia and Plantae, until 1960s – Modern classification • major categories: domain, kingdom, phylum, class, order, family, genus, and species • domains: Archaea, Eubacteria, Eukarya • kingdoms: Eukarya contains 3 kingdoms, Fungi, Plantae and Animalia • protists: eukaryotic organisms that do not fit the definition of animal, plant or fungus • Phylogeny: – phylogenetic tree – phenetics – cladistics