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Fig. 8.26 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Cerebrum Third neuron in pathway Thalamus Midbrain Pons Second neuron in pathway Medulla oblongata Receptor First neuron in pathway Dorsal column Spinal cord Fig. 8.27 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Lower limb Trunk Primary somatic sensory cortex Upper limb Somatic sensory association area Primary motor cortex Premotor area Prefrontal area Central sulcus Head Sensory speech area (Wernicke area) Motor speech area (Broca area) Visual cortex Visual association area Primary auditory cortex Taste area (beneath surface in insula) Auditory association area Lateral view Fig. 9.1 Copyright © McGraw-Hill Education. Permission required for reproduction or display. SENSES Receptors distributed over a large part of the body Receptors localized within specific organs General senses Located in skin, muscles, joints Special senses Located in internal organs Somatic Visceral Smell Touch Pressure Pain Balance Taste Temperature Hearing Sight Proprioception Pain Pressure Fig. 9.2 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Hair Free nerve endings (respond to painful stimuli, temperature, itch or movement) Merkel disks (detect light touch and superficial pressure) Epidermis Meissner corpuscles (involved in fine, discriminative touch) Dermis Hair follicle receptor (detects light touch) Ruffini end organ (detects continuous touch or pressure) Pacinian corpuscle (detects deep pressure, vibration, and position) Fig. 8.18 Copyright © McGraw-Hill Education. Permission required for reproduction or display. To brain From brain Sensory neuron 3 2 Quadriceps femoris muscle (extensor) Sensory receptor 4 Motor neuron 1 Patellar tendon Hammer tap Patellar ligament Flexor muscles Neuromuscular junction Knee-jerk reflex 1 Sensory receptors in the muscle detect stretch of the muscle. 2 Sensory neurons conduct action potentials to the spinal cord. 3 Sensory neurons synapse with motor neurons. Descending neurons (black) within the spinal cord also synapse with the neurons of the stretch reflex and modulate their activity. 4 Stimulation of the motor neurons causes the muscle to contract and resist being stretched. Fig. 9.2 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Hair Free nerve endings (respond to painful stimuli, temperature, itch or movement) Merkel disks (detect light touch and superficial pressure) Epidermis Meissner corpuscles (involved in fine, discriminative touch) Dermis Hair follicle receptor (detects light touch) Ruffini end organ (detects continuous touch or pressure) Pacinian corpuscle (detects deep pressure, vibration, and position) Fig. 9.3 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Lungs and diaphragm Liver and gallbladder Heart Esophagus Liver and gallbladder Stomach Kidney Colon Appendix Urinary bladder Ureter Fig. 9.4 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Olfactorybulb Frontal bone Olfactory tract Fibers of olfactory nerve Cribriform plate of ethmoid bone Nasal cavity Palate Nasopharynx (a) Interneurons Olfactory tract Olfactory bulb Foramen Cribriform plate Connective tissue Olfactory epithelium Mucous layer on epithelial surface (b) Olfactory nerve Axon Olfactory neuron Dendrite Cilia Fig. 8.37 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Anterior Olfactory bulb (olfactory nerves [I] enter bulb) Optic nerve (II) Olfactory tract Oculomotor nerve (III) Trochlear nerve (IV) Optic chiasm Trigeminal nerve (V) Pituitary gland Abducens nerve (VI) Mammillary body Facial nerve (VII) Pons Vestibulocochlear nerve (VIII) Glossopharyngeal nerve (IX) Vagus nerve (X) Medulla oblongata Hypoglossal nerve (XII) Accessory nerve (XI) Posterior Inferior view Fig. 8.27 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Lower limb Trunk Primary somatic sensory cortex Upper limb Somatic sensory association area Primary motor cortex Premotor area Prefrontal area Central sulcus Head Sensory speech area (Wernicke area) Motor speech area (Broca area) Visual cortex Visual association area Primary auditory cortex Taste area (beneath surface in insula) Auditory association area Lateral view Fig. 9.5 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Root of tongue Epiglottis Palatine tonsil Papillae (a) Epithelium Papilla Taste bud (b) Supporting cell Taste cell Taste hair (c) Nerve fiber of sensory neuron Taste pore Epithelium Fig. 8.37 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Anterior Olfactory bulb (olfactory nerves [I] enter bulb) Optic nerve (II) Olfactory tract Oculomotor nerve (III) Trochlear nerve (IV) Optic chiasm Trigeminal nerve (V) Pituitary gland Abducens nerve (VI) Mammillary body Facial nerve (VII) Pons Vestibulocochlear nerve (VIII) Glossopharyngeal nerve (IX) Vagus nerve (X) Medulla oblongata Hypoglossal nerve (XII) Accessory nerve (XI) Posterior Inferior view Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.6 Taste area of cortex Thalamus Nucleus of tractus solitarius V Chorda tympani VII IX X Foramen magnum Facial nerve (VII) Trigeminal nerve (V) (lingual branch) Glossopharyngeal nerve (IX) Vagus nerve (X) Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.17 External ear Middle ear Inner ear Auricle Copyright © McGraw-Hill Education. Permission required for reproduction or display. Temporal bone Tympanic membrane External auditory canal Chorda tympani Semicircular canals Oval window Vestibulocochlear nerve Cochlear nerve Vestibule Cochlea Round window Auditory tube Malleus Incus Stapes Auditory ossicles in the middle ear Frontal Section Fig. 9.21 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Otoliths Gelatinous mass (d) Microvilli Utricular macula Saccular macula Nerve fibers of vestibular nerve Vestibule Utricle Saccule (a) Part of macula Hair cell Supporting cells (b) ©Susumu Nishinag/Science Source (c) Fig. 9.22 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Endolymph in utricle Gelatinous mass Hair cell Supporting cell Force of gravity Macula Vestibular nerve fibers (a) (b) (all): ©Trent Stephens Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.17 External ear Middle ear Inner ear Auricle Copyright © McGraw-Hill Education. Permission required for reproduction or display. Temporal bone Tympanic membrane External auditory canal Chorda tympani Semicircular canals Oval window Vestibulocochlear nerve Cochlear nerve Vestibule Cochlea Round window Auditory tube Malleus Incus Stapes Auditory ossicles in the middle ear Frontal Section Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.23 Semicircular canals Ampullae Vestibular nerve Cupula (a) Microvilli Cupula Crista ampullaris (b) Hair cell Nerve fibers to vestibular (c) nerve Fig. 9.24 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Endolymph causes movement of cupula. Cupula Endolymph in semicircular canal Hair cell Crista ampullaris Movement of semicircular canal with body movement (b) (a) ©Julie Jacobson/AP Images Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.17 External ear Middle ear Inner ear Auricle Copyright © McGraw-Hill Education. Permission required for reproduction or display. Temporal bone Tympanic membrane External auditory canal Chorda tympani Semicircular canals Oval window Vestibulocochlear nerve Cochlear nerve Vestibule Cochlea Round window Auditory tube Malleus Incus Stapes Auditory ossicles in the middle ear Frontal Section Fig. 9.18 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Periosteum of bone (inner lining of bony labyrinth) Semicircular canals Cochlear Scalavestibuli (filled with perilymph) Membranous labyrinth Copyright ©nerve McGraw-Hill Education. Permission required for reproduction or display. Vestibular membrane Tectorial membrane Cochlear duct (filled with endolymph) Vestibule Oval window Spiral ligament Cochlea Basilar membrane Round window Scalatympani (filled with perilymph) Spiral lamina Apex Cochlearganglion (a) (b) Cochlear duct Vestibular membrane Tectorial membrane Microvilli Basilar membrane Cochlear nerve Supporting cells Hair cell Spiral lamina Hair cell Nerve endings of cochlear nerve (c) (d) Courtesy of A. J. Hudspeth (e) Spiral organ Spiral ligament Fig. 9.19 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Oval window Stapes Incus Cochlear nerve Malleus Scala vestibuli Tympanic membrane Scala tympani 3 Cochlear duct (contains endolymph) 2 External auditory canal Space between bony labyrinth and membranous labyrinth (contains perilymph) 4 1 5 Basilar membrane 7 Round window Auditory tube 1 Sound waves strike the tympanic membrane and cause it to vibrate. 2 Vibration of the tympanic membrane causes the malleus, the incus, and the stapes to vibrate. 3 The base of the stapes vibrates in the oval window. 4 Vibration of the base of the stapes causes the perilymph in the scala Vestibular membrane 6 Membranous labyrinth Tectorial membrane Spiral organ 6 Vibration of the endolymph causes displacement of the basilar membrane. Short waves (high pitch) cause displacement of the basilar membrane near the oval window, and longer waves (low pitch) cause displacement of the basilar membrane some distance from the oval window. Movement of the basilar membrane is detected in the hair cells of the spiral organ, which are attached to the basilar membrane. Vibrations of the perilymph in the scala vestibuli and of the basilar membrane are transferred to the perilymph of the scala tympani. vestibuli to vibrate. 7 Vibrations in the perilymph of the scala tympani are transferred to the 5 Vibration of the perilymph causes the vestibular membrane to vibrate, which causes vibrations in the endolymph. round window, where they are dampened. Fig. 9.20 Copyright © McGraw-Hill Education. Permission required for reproduction or display. 1 Sensory axons from the cochlear ganglion terminate in the cochlear nucleus in the brainstem. 2 Axons from the neurons in the cochlear nucleus project to other brainstem nuclei or to the inferior colliculus. 3 Axons from the inferior colliculus project to the thalamus. 4 Thalamic neurons project to the auditory cortex. Thalamus 4 Auditory cortex Auditory cortex 3 2 Cochlear ganglion 2 Inferior colliculus Vestibulocochlear nerve Other brainstem nucleus 1 Cochlear nucleus Nerve to stapedius Frontal section Fig. 9.7 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Upper eyelid Eyebrow Eyebrow Pupil Iris Lateral angle of eye Lower eyelid Medial angle of eye (a) Muscle to upper eyelid Superior rectus muscle Conjunctiva over eye Conjunctiva of eyelid Sebaceous gland Upper eyelid Connective tissue plate Lacrimal gland Cornea Eyelash Lacrimal canaliculi Lacrimal ducts Lacrimal sac Skin Loose connective tissue Nasolacrimal duct (c) Orbicularis oculi muscle Inferior rectus muscle Orbicularis oculi muscle Inferior oblique muscle Conjunctiva of eyelid (b) ©McGraw-Hill Education/Eric Wise Connective tissue plate Lower eyelid Fig. 9.9 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Vitreous chamber (filled with vitreous humor) Vitreous humor Optic nerve Sclera Cornea Fibrous tunic Suspensory ligaments Anterior chamber Posterior chamber Both filled with aqueous humor Pupil Lens Retina (nervous tunic) Iris Ciliary body Choroid Vascular tunic Fig. 9.14 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Medial Lateral Optic disc Macula Retinal vessels Fovea centralis (a) (b) ©Steve Allen/Getty Images RF Fig. 9.15 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Near vision Distant vision Ciliary muscles in the ciliary body are relaxed. Ciliary muscles in the ciliary body contract, moving ciliary body toward lens. Tension in suspensory ligaments is high. Tension in suspensory ligaments is low. A FP FP A Lens flattened Lens thickened (a) (b) Fig. 9.10 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Ciliary muscles of the ciliary body Lens Suspensory ligaments Fig. 9.11 Copyright © McGraw-Hill Education. Permission required for reproduction or display. Circular smooth muscles (a) Radial smooth muscles (b) Fig. 9.8 Copyright © McGraw-Hill Education. Permission required for reproduction or display. View Superior Muscleto uppereyelid (cut) Superior rectus Superior oblique Optic nerve Inferior rectus Lateral rectus Inferior oblique Inferior Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.12 Choroid Pigment cell Pigmented retina Photoreceptor layer Cone cell Direction of action potential propagation Rod cell Horizontal cell Sensory retina Bipolar cell Interneurons Direction of light Ganglion cell Nerve fibers to optic nerve (a) Optic nerve Disc Outer membrane Disc Outer segment Folding of outer membrane to form discs Rod Cone Disc Inner segment (e) (b) Nuclei Outside of disc membrane Disc membrane Axons Synaptic ending Rod Inside of disc membrane (c) (d) Cone (f) ©Steve Gschmeissner/Science Source Opsin Retinal Rhodopsin Fig. 9.13 Copyright © McGraw-Hill Education. Permission required for reproduction or display. 1 Opsin Retinal Rhodopsin Light 1 Rhodopsin is composed of opsin and retinal. 2 6 2 Light causes retinal to change shape, which DIGESTIVE activates rhodopsin. 3 Tissue damage to intestinal lining and liver as a result of decreased blood flow; bacteria of intestines may cause systemic infection; liver releases blood-clotting Activated rhodopsin factors in response to injury. stimulates cell changes that result in vision. URINARY 4 Following rhodopsin Urine production decreases in activation, retinal detaches response to low blood from opsin. volume; tissue damage to 3 Retinal kidneys due to low blood flow. 5 Energy from ATP is required to bring retinal back to its original form. Cell changes that result in vision 5 Energy (ATP) 6 Retinal recombines with opsin to form rhodopsin (return to step 1). 4 Retinal Fig. 9A Copyright © McGraw-Hill Education. Permission required for reproduction or display. (a) (b) (all): ©Prisma Bildagentur AG/Alamy Copyright © McGraw-Hill Education. Permission required for reproduction or display. Fig. 9.16 Nasal parts of visual fields 1 1 Each visual field is divided into temporal Temporal Temporal part of left part of right and nasal Copyright halves. © McGraw-Hill Education. Permission required for reproduction or display. visual field visual field 2 After passing through the lens, light from each half of a visual field projects to the Lens opposite side of the retina stimulating 2 Left eye receptors. 3 Axons from the retina pass through the optic nerve to the optic chiasma, where some cross. Axons from the nasal retina cross, and those from the temporal retina do not. Temporal retina (lateral part) Nasal retina (medial part) 3 Optic nerves 4 Superior colliculi Optic chiasm 4 Optic tracts extend from the optic chiasm (with or without crossing) to the thalamus. Collateral branches of the axons in the optic tract synapse in the superior colliculi of the midbrain. Optic tracts Thalamus 5 Optic radiations extend from the thalamus to the visual cortex of the occipital lobe. Optic radiations 6 The right part of each visual field (dark green and light blue) projects to the left side of the brain, and the left part of each visual field (light green and dark blue) projects to the right side of the brain. Visual cortex in each hemisphere 5 6 Occipital lobe (a) Optic nerve Left monocular vision Binocular vision Optic chiasm Optic tract Thalamus Optic radiation Visual cortex (b) (c) ©McGraw-Hill Education/Rebecca Gray, photographer/Don Kincaid, dissections Right monocular vision