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David Sadava, David M. Hillis, H. Craig Heller, May R. Berenbaum La nuova biologia.blu Le cellule e i viventi Community and Population Ecology What Are Ecological Communities? An ecological community consists of all the species that coexist and interact in a defined area. What Are Ecological Communities? A food chain depicts who eats whom in a community. Food chains are interconnected to make a more realistic depiction of community feeding relationships, called a food web. What Are Ecological Communities? An organism’s trophic level indicates where in the food chain it obtains its energy. What Are Ecological Communities? Communities are characterized by: 1. Species composition — the number and kinds of species they contain. 2. The relative abundances of those species. Communities can be characterized by the distribution of energy and biomass within them. What Are Ecological Communities? At each trophic level energy is lost to metabolism and respiration. Only 10% of energy of one trophic level is transferred to the next level. Energy and Biomass Distributions Ecological efficiency: overall transfer of energy from one trophic level to the next (ratio of consumer production to producer production). Pyramid diagrams show energy, biomass or numbers of individuals at each trophic level. What Are Ecological Communities? Gross primary productivity (GPP) is the rate at which primary producers turn solar energy into stored chemical energy via photosynthesis. Net primary productivity (NPP) is the rate at which energy is incorporated into primary producers by growth and reproduction. NPP = GPP – R R = energy lost through respiration How Does Energy Flow Through the Global Ecosystem? Different types of ecosystems have different rates of production. Geographic Variation in Terrestrial and Marine NPP How Do Specific Nutrients Cycle through the Global Ecosystem? The patterns of movements of elements around the planet are called biogeochemical cycles. Properties of biogeochemical cycles depend on the physical and chemical nature of each element and how it is used by organisms. Biogeochemical Cycles: Hydrologic Cycle The hydrologic cycle is the cycling of water through the global ecosystem. The sun powers the hydrologic cycle by causing evaporation from the ocean surfaces. More water is evaporated from the oceans than is returned by precipitation. The excess falls as precipitation on land. Biogeochemical Cycles: Nitrogen Cycle N2 gas makes up 78% of the atmosphere, but most organisms cannot use N in this form. Some bacteria can convert N2 into a form usable by plants (nitrogen fixation). Other microorganisms carry out denitrification, returning N2 to the atmosphere. Biogeochemical Cycles: Carbon cycle CO2 is taken up from the atmosphere by autotrophs and incorporated into organic molecules by photosynthesis. Carbon is returned to the atmosphere by the metabolism of organisms. Biogeochemical Cycles: Carbon cycle CO2 is being released into the atmosphere faster than it can be dissolved in oceans and taken up by organisms. Biogeochemical Cycles: Phosphorus cycle Phosphorus is an essential nutrient for DNA, RNA, ATP, and phospholipids. It lacks a gaseous phase. There is very little in the atmosphere. Phosphorus cycles rapidly in organisms, but very slowly in Earth’s crust. Biogeochemical Cycles: Sulfur cycle Emissions of SO2 and H2S from volcanoes and fumaroles are the only non-microbial sources of S in the atmosphere. Microbial decomposition in marine and terrestrial environments returns S to the atmosphere. Burning fossil fuels contributes SO2 to the atmosphere, where it can form sulfuric acid (H2SO4). Acidification Reduces Fish Species Richness Sulfuric and nitric acid (HNO3) form acid precipitation. How Do Interactions among Species Influence Communities? Habitat: the space where organisms live. Ecosystem engineers: organisms that influence the ecosystem structure. A species that exerts influence on a community that is out of proportion with its abundance is called a keystone species. They may influence number of species and the number of trophic levels. How Do Interactions among Species Influence Communities? A species’ niche is the set of physical and biological conditions it needs to survive, grow, and reproduce. Competitive exclusion principle: no two species can share the same limiting resource indefinitely. How Do Disturbances Affect Ecological Communities? Patterns of change in community composition following disturbance is called succession. Primary succession begins after a disturbance that removes all organisms and exposes bare substrate. How Do Disturbances Affect Ecological Communities? Secondary succession occurs following disturbances in which some organisms survive. How Do Ecologists Measure Populations? A population consists of individuals of a species that interact with one another within a given area at a particular time. To study populations, ecologists count individuals, determine their ages, and calculate the rates at which individuals enter and leave the population. The number of individuals per unit area or volume is the population density. Dispersion Patterns Dispersion: distribution of individuals in space. Dispersion determines patterns of interaction among individuals. How Do Ecologists Measure Populations? The study of population processes is known as demography. Demographic events: births, deaths, immigration, and emigration. How Do Ecologists Measure Populations? To study populations, ecologists count individuals, determine their ages, and calculate the rates at which individuals enter and leave the population. Individuals are often tagged or marked in some way to facilitate counting. How Do Ecologists Measure Populations? Population sizes are often estimated from representative samples using statistical methods. Individuals may be counted within measured areas called quadrats and plants are often counted along a linear transect. The mark–recapture method involves capture, marking, and releasing some individuals, then later capturing another sample of individuals. What Factors Limit Population Densities? All populations have the potential for exponential growth. Carrying capacity (K) is the number of individuals of a species that can be supported in an environment indefinitely. This pattern is called logistic growth— S-shaped curve. What Factors Limit Population Densities? Some species achieve higher population densities than others. Three factors influence this: • Abundance of resources • Body size • Complex social organization. Density-dependent regulation factors. Density-independent factors. Two Life History Strategies What Types of Interactions Do Ecologists Study? Interactions among species can be placed into five broad categories. How Do Antagonistic Interactions Evolve? Prey species have many defenses, including running away or staying motionless to avoid detection. Morphological defenses include shells, tough skin, spines, and hair. Chemical defenses are common in small or weak prey and sessile organisms. How Do Antagonistic Interactions Evolve? Prey species that are toxic may advertise the fact with bright coloration or warning signs: aposematism or warning coloration. Some prey species are mimics. Adapted from Life: The Science of Biology, Tenth Edition, Sinauer Associates, Sunderland, MA, 2014 Inc. All rights reserved