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Unit 10: Evolution Ch. 29 Evolution History of Earth • Earth ~4.6 billion years old – early Earth • hostile environment – volcanoes, lightning, meteorites, UV light • primitive atmosphere – methane, ammonia, hydrogen, water vapor » What’s missing? Origin of Life • Life ~3.5 billion years old • Miller & Urey – simulated primitive atmosphere • zapped w/ electricity – ~1 week amino acids & other simple organic molecules formed (in water) » precursors to 1st cells The First Organisms: Heterotrophs • “Heterotroph Hypothesis” or “Primordial Soup Hypothesis” – Started w/ simple compounds complex compounds primitive pre-cells prokaryotes (1st cells) • 1st cells were heterotrophs – Why heterotrophic? » b/c no O2 for cellular respiration/photosynthesis – fed on organic material (primordial soup in ocean) » Similar to some present-day bacteria, obtained energy from simple chemical reactions Later Organisms: Autotrophs • prokaryotic autotrophs followed heterotrophs – What did they release into the atmosphere? • oxygen (causing the oxygen revolution!) – Why is this so important? » allowed development of eukaryotes & complex organisms stromatolites What is Evolution? • theory that species have changed gradually over time – individual’s genes stay the same – population’s gene pool (total genes) may change • evolves as allele frequencies (%) (how often an allele occurs) change due to natural selection Mechanisms of Evolution • • • • migration genetic drift mutation natural selection Mechanisms of Evolution • migration – gene frequencies change as individuals move in or out of the population • genetic drift – change in gene pool of a small population that takes place strictly by CHANCE Mechanisms of Evolution • mutation – random change in the DNA sequence of a gene • can change amino acid sequence & protein coded for – can change the way the trait is expressed • not all mutations matter in evolution – must be heritable (in gametes) Mechanisms of Evolution • natural selection – organisms with variation best suited for environment tend to survive, & reproduce (“survival of the fittest”) • more offspring will have favorable adaptation than before – acts on individuals, but causes evolution of population •Which trait was a better adaptation and selected? •How were frequencies of different alleles affected? Explanations for How Evolution Happens • Jean Baptist Lamarck – “Theory of Acquired Characteristics” • acquired traits are passed on – leads to changes in species –experimentation failed to support Lamarckism Explanations for How Evolution Happens • Charles Darwin – naturalist on HMS Beagle • collected fossils & specimens • observed tremendous biodiversity Climate varied from island to island. intermediate shell domed Shell large rainfall saddlebacked shell desert Explanations for Evolution – concluded, life forms can & do change…. • but, questioned how this change occurred… –concluded evolution or “descent with modification” »still needed to test ideas before explaining how or why such changes happen Darwin’s Explanation: Natural Selection – wrote On the Origin of Species by Means of Natural Selection (1859) – Darwin’s Main Points: 1. Overproduction of offspring 2. Struggle to survive – competition 3. Variation within population 4. Successful reproduction – “survival of the fittest”/best adapted organisms are “naturally selected” Video: Who Was Charles Darwin? Darwin’s Explanation: Natural Selection 1. Overproduction of offspring – Why is this necessary? • to ensure that at least some will survive long enough to reproduce & pass on genes Video: Darwin & Malthus Darwin’s Explanation: Natural Selection 2. Struggle to survive – more organisms than resources to go around leads to competition • For what? – food, shelter, mates, space, light, water, nutrients, etc. • Survivors – Who are they? – How did they get these traits? – Were these traits a response to their environment? Darwin’s Explanation: Natural Selection 3. variation w/in population – What causes variety? • Sexual reproduction • Crossing over during meiosis • Arrangement of chromosomes during meiosis • Mutations – Darwin did NOT have knowledge of genetics Darwin’s Explanation: Natural Selection • variations – can be adaptations – are NOT a response to environment • just happen to be “useful” when environment changes camouflage Darwin’s Explanation: Natural Selection 4. successful reproduction – natural selection • Those organisms best suited for environment tend to survive, reproduce, & pass on genes (“survival of fittest”) – more offspring have favorable adaptation than before Pepper Moth “Game” http://www.techapps.net/interactives/pepperMoths.swf Principles of Natural Selection "Interactive" How would Darwin adjust the description given by Lamarck about the length of giraffes necks? Lamarck vs. Darwin Darwin’s Explanation: Wallace’s Same Conclusion • Alfred Wallace (1858) – arrived at same basic explanation as Darwin • Darwin usually given credit – but BOTH men should be credited Evidence to support the Theory of Evolution • indirect evidence – fossil record – comparative anatomy – comparative embryology – comparative biochemistry • direct evidence – current observations Indirect Evidence: Fossil Record • Fossil Record – Evidence of: • change in organisms over time • age of the Earth – Limited b/c: • • decomposition type of material – • incomplete record – • What fossilizes best? Why? easily disrupted – Why? Which layer is youngest? oldest? Explain… Law of Superposition Indirect Evidence: Comparative Anatomy • comparative anatomy – compares structures of diff. organisms living today 1. homologous structures 2. analogous structures 3. vestigial structures Indirect Evidence: Comparative Anatomy 1. homologous structures • similar structure w/ modifications − evidence of common ancestor • becoming more different (divergent evolution) Indirect Evidence: Comparative Anatomy • 2. analogous structures – similar function, but not structure • due to similar environment/use • do NOT have common ancestor – becoming more similar (convergent evolution) Indirect Evidence: Comparative Anatomy • 3. vestigial structures – structures with reduced size or function • often no apparent use anymore – evidence that organism has changed Indirect Evidence: Comparative Embryology • embryonic development – early stages very similar • later stages more distinction – inherited same basic body plan from common ancestor • greater # similar stages more closely related Indirect Evidence: Comparative Biochemistry • similar organic compounds coded for by similar DNA – ex. proteins Indirect Evidence: Comparative Biochemistry • DNA most reliable form of evidence • What is the significance of 2 very different species sharing large amounts of the same DNA? – evidence of common ancestry • higher %, more closely related – human & chimpanzees 98% similar – humans & other mammals 80% similar Direct Evidence: Observation • “rapid” evolution 1. antibioticresistant bacteria 2. pesticideresistant insects