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Chapter 5: Interactions: Environments and Organisms Ecological Concepts (Figure 5.1) Ecology: The study of ways of organisms interacts with each other and with their surroundings. Environment: Everything that affects and organism during its lifetime is collectively known as its environment, which contains abiotic (nonliving) and biotic (living) factors. Abiotic Factors: Nonliving things that influence an organism, including energy, nonliving matter, living space, and process that involves the interactions of nonliving matter and energy. Energy: Required by all organisms to maintain themselves. Ultimate source: Sun. Plants: Directly use sun by photosynthesis of sugar. Animals: Obtain energy by eating plant. Matter: Provide the structural framework of organisms, which obtain these materials from environment. Atoms: In form of matter, which become part of an organism’s body structure for a short period and eventually all of them are returned to the environment through respiration, excretion, or death and decay. Space: The place organisms inhabit that has a particular structure and location. Ex. Ocean, mountains, near equator area, near poles area. Ecological processes: Involving interaction between matter and energy. Ex. Solar radiation affects ocean currents, generates wind and climate change, evaporation of water into atmosphere and fall as precipitation. Biotic Factors: Include all forms of life with which it interacts. Ex. Plants: photosynthesis Animals: eat other organisms Bacteria and fungi: cause decay Bacteria, viruses, parasitic organisms: cause disease Limiting Factors: A shortage or absence of a factor can restrict the success of the species is known as a limiting factor, which may be abiotic or biotic and can be quite different form one species to the other species. Ex. Plants: scarcity of water, light or specific soil nutrients. Animals: climate or availability of a specific food. Aquatic Species: cool oxygenated water. Range of Tolerance: each species has its own. Ex. Trout: unable to survive in low [O2] and high temperature. 1 Habitat and Niche Habitat: The place an organism lives Ex. Moss: cool, moist and shady. Niche: The functional role an organism has in its surroundings. Ex. Beaver: building dam providing deep water for protection and habitat for ducks and fish, flooding area, killing trees, become victim when animal attracts to pond, eating suitable woody trees. When foods are gone, they migrate to other area along the stream. The Role of Natural Selection and Evolution General rule: Organisms are well adapted to their surroundings and fill a particular niche. What leads to this high adaptation? Genes: The distinct pieces of DNA that determines the characteristics an individual displays. Population: Organisms of the same kind found within a specific geographic region. Individuals will have very similar sets of genes but there are some variations. Genes are passed from one generation to the next by reproduction. Species: A population of all the organisms potentially capable of reproducing naturally among themselves or interbreeding and having fertile offspring. (Fig. 5.8) Usually requires specific conditions. Ex. Horse-horse, donkey-donkey, horse-donkey (= mule, infertile) Asexual reproduction: 1) Organisms produce copies of them without mating. (non-species) 2) Organisms produce copies of them by mating (species) and without mating (non-species). Natural Selection by Charles Darwin: The mechanism causes evolution to occur. Not understanding the concepts of genes but understanding characteristics were passed from parents to offspring. Observed highly adaptive nature of the relationship between organisms and their environment and developed the concept of natural selection to explain how this adaptation came about. The process that determines which individuals within a species will reproduce and pass their genes to the next generation. Evolution: The changes we seen in the genes and the characteristics displayed by successive generations of a population of organisms over time. Steps involved: 1) Genetic Variation: Individuals within a species show genetically determined variation; some of the variations are useful and other are not. (Ex. color variation: white, yellow, black, red). (Fig. 5.10) 2 2) Excessive Reproduction: Organisms within a species typically produce many more offspring than are needed to replace the parents when they die. Most of the offspring die. Ex. Blueberries, rabbits. (Fig. 5.11) 3) The excessive number of individuals results in a shortage of specific resources. Ex. Individual species compete with each other for food, space, and mate that are in limited supply. 4) Because of variation among individuals, some have a greater chance of obtaining needed resources and, therefore, have a greater likelihood of surviving and reproducing than others. Ex. Individuals that have genes that allow them to obtain needed resources and avoid threats to their survival will be more likely to survive and reproduce. 5) As time passes and each generation is subjected to the same process of natural selection, the percentage of individuals showing favorable variations will increase and those having unfavorable variations will decrease. Evolutionary Patterns Speciation: 1) The production of new species from previously existing species. 2) It is thought to occur as a result of a species dividing into two isolated subpopulations. 3) If two subpopulation contain some genetic differences and their environments are somewhat different, natural selection will work on the two groups differently and they wil begin to diverge from each other. 4) Eventually, the differences may be so great that the two subpopulations are not able to interbreed. At this point, they are two different species. (Fig. 5.13) 5) For plants, polyploidy is another common mechanism results in new species. Polyploidy is a condition in which the number of sets of chromosomes in the cells of the plant is increased. Extinction: 1) The loss of an entire species and is a common feature of the evolution of organisms. 2) The environment in which organisms exist does not remain constant over long time periods. 3) Those species lack the genetic resources to cope with a changing environment go extinct. 4) Humans have had a significant impact on the extinction of many kinds of species. Coevolution: 1) The concept that two or more species of organisms can reciprocally influence the evolution of other organisms. Ex. Tubular flower (nectar at the base of flower) and hummingbird (with long bill) Kinds of Organism Interactions Predation: A common interaction occurs when one organism, known as predator, kills and eats another, known as the pray. Ex. Lion and zebra, robin and earthworm, wolf and moose, etc. A high reproductive rate is an adaptation common to many pray species. Competition: Two organisms strive to obtain the same limited resource. 3 1) Intraspecific competition: Competition between members of the species. Ex. Intraspecific competition for sunlight among pine trees results in the tall, straight trunks. (Fig. 5.17) 2) Interspecific competition: Competition between members of different species. Ex. Hawks, foxes, owls, and coyotes may compete the same prays. Competition and Natural Selection 1) The major force in shaping the evolution of a species is the competition among members of the same species. 2) When resources are limited, the less well-adapted individuals are more likely to die or be denied mating privileges. 3) Because of the most successful organisms are likely to have larger numbers of offspring, each succeeding generation will contain more of the genetic characteristics that are favorable for survival of the species in that particular environment. Symbiotic Relationships Symbiosis: A close, long-lasting, physical relationship between two different species. Categories: 1) Parasitism: (Fig. 5.20) A relationship in which one organism, known as the parasite, lives in or on another organism, known as host, from which it derives nourishment. Parasites, usually much smaller than the host, harm the host and gradually kills the host. o Ectoparasites: live on the surface of their hosts. Ex. Fleas, lice, etc. o Endoparasites: live inside the bodies of their hosts. Ex. Tapeworm. 2) Commensalism: A relationship between organisms in which one organism benefits while the other is not affected. It’s possible tovisualize a parasitic relationship evolving into a commensal one. Ex. Shark and remora. 3) Mutualism: Another kind of symbiotic relationship and is actually beneficial to both species involved. In many, the relationship is obligatory: one can not live without the other. In others, the species can exist separately but are more successful when they are involved in the relationship. Ex. Fungus and root association (i.e. mycorrhizae): fungi obtain nutrients like PO43- and NO3- to make p and N available to plants. Some Relationships are Difficult to Categorize Ex. Nest (Brood) parasitism: (Fig. 5.23) Cowbirds and European cuckoos laid eggs in other birds’ nests and let other birds to raise their young. Community and Ecosystem Interactions Community: An assemblage of all the interacting populations of different species of organisms in an area. Some plays minor roles, while others play major roles. 4 Ecosystem: A define space in which interactions take place between a community, with all its complex interrelationship and the physical environment. The physical world has a major impact on what kinds of plants and animals can live in an area. Major Roles of Organisms in Ecosystems 1) Producers: Organisms that able to use sources of energy to make complex, organic molecules from simple inorganic substances in their environment. Ex. Plants, algae, phytoplankton, etc. 2) Consumers: Organisms that require organic matters as a source of food to provide themselves with energy and the organic molecules necessary to build their own bodies. Primary consumers (i.e. herbivores): animals that eat producers as a source of food. (Cows, horses, etc.) Secondary consumers (i.e. carnivores): animals that eat other animals. (Eagles, lions, etc.) Omnivores: Animals that eat both plants and animals. (Foxes, bears, etc.) Decomposers: Organisms that use nonliving organic matter as a source of energy and raw materials to build their bodies. (Bacteria, fungi, small animals, etc.) Keystone species: One that has a critical role to play in the maintenance of specific ecosystems. (Fig. 5.24) Energy Flow through Ecosystem (Fig. 5.26) Ecosystem: 1) A stable, self-regulating unit via growing, reproducing, dying, and decaying. 2) Having a constant input of energy to retain its stability. 3) Sunlight: The only significant input of energy. 4) Producers trap the solar energy through photosynthesis to make it available to the ecosystem. 5) Energy stored in the producers are transferred to other organisms when the producers are eaten. Tropic Levels: Each level in the flow of energy through an ecosystem. (Fig. 5.25) 1) First tropic level: producers. 2) Second tropic level: herbivores. 3) Third tropic level: carnivores that eat herbivores. 4) Fourth tropic level: carnivores that eat other carnivores. Energy Relationships: 1) 2nd law of thermodynamics: When energy is converted from one form to another, some of energy is converted to a non-useful form, typically low-quality heat, which is dissipated to the surroundings and warms the air, water or soil. 2) The amount of energy contained in higher tropic levels is considerably less than that at lower level, as it requires energy to chew, defend nests, or produce and raise offspring. 3) Ecologists use biomass (the weight of mass in a tropic level) to measure the amount of energy contained in each tropic level. 5 Food Chain and Food Web Food chain: A series of organisms occupying different tropic levels through which energy passes as a result of one organism consuming another. (Fig. 5.27) Detritus: The small bits of nonliving organic material. Food web: When several food chains overlap and intersect. (Fig. 5.28) Nutrient Cycles in Ecosystems----Biogeochemical Cycle Biogeochemical cycle: The nutrient cycle as all matter is made up of atoms and atoms are cycled between the living and nonliving portions of ecosystem. Carbon Cycle ((Fig. 5.29): The processes and pathways involved in capturing inorganic carbon-containing molecules, converting them into organic molecules that are used by organisms, and the ultimate release of inorganic carbon molecules back to the abiotic environment, for all living things are composed of organic molecules that contain atoms of carbon element. The role of producers: 6CO2 + 6H2O C6H12O6 + 6O2 The role of consumers: Herbivores eat plants or algae to break down complex organic molecules, which can be reassembled into the specific organic molecules that are part of its chemical structure. The carbon atom, which was once part of organic molecule in a producer, is now part of an organic molecule in an herbivores. During the process of respiration, the oxygen in atmosphere is used to break down large organic molecules into CO2 and H2O. Energy stored in chemical bonds is released by respiration and is lost as heat. By the same token, the same processes occur when carnivore eats herbivore. The role of decomposers: Decay process of decomposers involves respiration and releases CO2 and H2O back to the nature through recycling. The role of fossil fuels: Coal, oil, and natural gas are part of the carbon cycle. Organisms were buried and the organic compounds in their bodies were modified by geologic forces. The C atoms present in the fossil fuels were removed temporarily from the active, short-term C cycle. When fossil fuel are burned, the C reenters the active C cycle. Nitrogen Cycle: The cycling of N atoms between the abiotic and biotic components and among the organisms in an ecosystem. Very Few organisms are able to use tightly bonded N2. It must be in the forms of NO3- or NH3. The Role of Nitrogen-Fixing Bacteria: 1) Able to convert N2 that enters the soil into NH3. 2) Free-living nitrogen-fixing bacteria. 3) Symbiotic nitrogen-fixing bacteria: Having mutualistic relationship with certain plants (peas, beans, clover, etc) and certain trees (alders, etc.) 6 The Role of Producers and Consumers: 1) Construct proteins, DNA, and other important nitrogen-containing compounds. 2) When herbivores eat plants, the plant protein molecules are broken down to amino acid, a smaller building block. 3) Amino acids are reassembled to form proteins typical for the herbivores. 4) Same process is repeated throughout the food chain. The Role of Decomposers and Other Bacteria: 1) Involve in nitrogen cycle by breaking down nitrogen-containing organic molecules, releasing NH3, which can be directly used by many plants. 2) Nitrifying bacteria converts NH3 to NO2-, which can be converted to NO3- to be used by plants. 3) Denitrifying bacteria: Able to convert NO2- to N2 when O2 is absent. N2 is ultimately released into the atmosphere. Unique Features of the Nitrogen Cycle: (Fig. 5.30) Two significant differences from C cycle 1) Most of the difficult chemical conversions are made by bacteria and other microorganisms. 2) There is a secondary loop in the cycle that recycles nitrogen compounds directly from dead organisms and wastes directly back to producers. Agriculture and Nitrogen Cycle: 1) In naturally occurring soil, nitrogen is often a limiting factor for plant growth. 2) To increase yields, farmers provide extra sources of nitrogen in several ways. 3) Inorganic fertilizers are the primary method of increasing the nitrogen available. 4) Alternate planting between soybeans (nitrogen-yield crops) and corns (nitrogen-demanding crops). 5) Using composts from plants and manure from animals as fertilizers. Phosphorus Cycle (Fig. 5.31): 1) Phosphorus: A common element in plant structure. 2) Present in many important biological molecules: DNA, cell membrane, bones, and teeth. 3) Different from C and N cycles in one important respect: P is not present in the atmosphere as a gas. The ultimate source is rock, from which P is released by erosion and dissolves in water. Plants use dissolved P to construct the molecules they need. Animals obtain P by eating plants or other animals. Decomposers recycle P back into the soil by decomposing dead organisms or their wastes. Soluble phosphorus compounds are ultimately precipitated as deposits and geologic processes elevate these deposits and expose them to erosion, making deposits available to organisms. Waste products of animals often have significant amounts of phosphorus. 7 Human Impact on Nutrient Cycles Burning of Fossil Fuel: 1) Releasing large amounts of CO2, causing change of climate. 2) Seeking to reduce energy use and deforestation. 3) Altering the nitrogen cycle by doubling the amount of nitrogen available today when compared to pre-industrial times as heating O2 and N2 to high temperature to form more N-containing compounds. Conversion of Natural Ecosystem to Agriculture: 1) Forest, wetland, and grassland ecosystems tend to store carbon for long periods, while agricultural ecosystems store carbon only temporarily. 2) Converting to agricultural ecosystem has increased the amount of CO2 in the atmosphere (linking to global warming) has disrupted the natural C cycle. 3) Use of fertilizers, labeling in order of N, P, and K, to increase crops production has significantly altered several nutrients cycles. 4) C, N, P, K and other elements are cycled within ecosystems, as when crops are harvested, these elements are removed. 5) Farmers not only return N, P, and K, they also analyze for other less prominent elements and add them to their fertilizer mixture. Agricultural Runoff: 1) The nutrients in fertilizers are intended to become incorporated into the bodies of the plants and animals that we raise for food. 2) Applying too much N or P as fertilizer or if applying at the wrong time, much of this fertilizer is carried into aquatic ecosystems. 3) Raising large number of animals for food in concentrated settings results in huge amounts of animal wastes that contain N and P to enter local water sources. 4) The presence of large amount of N and P in fresher water or salt water results in increased rates of growth of bacteria, algae, and aquatic plants, which lead to low oxygen concentration and many organisms die. When decomposers use oxygen from the water as they break down the dead organic matter. 5) Many algae are toxic and when their numbers increase significantly, fish are killed and incidents of human poisoning occur. 6) Fertilizers use in the agricultural center of USA results in death of fish in the Gulf of Mexico. (Fig. 5.32) 8