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Invertebrates Section Outline Section 29-1 29–1 Invertebrate Evolution A.Origin of the Invertebrates 1.The First Multicellular Animals 2.Beginnings of Invertebrate Diversity B.Invertebrate Phylogeny C.Evolutionary Trends 1. Specialized Cells, Tissues, and Organs 2. Body Symmetry 3. Cephalization 4. Segmentation 5. Coelom Formation 6. Embryological Development Section 29-1 Compare/Contrast Comparing Invertebrates Major Characteristics Sponges Cnidarians Flatworms Roundworms Germ Layers Absent Two Three Three Body Symmetry Absent Radial Bilateral Bilateral Cephalization Absent Absent Present Present Coelom Absent Absent Absent Pseudocoelom Early Development — — Protostome Protostome Pg. 748-749 Section 29-1 Compare/Contrast Comparing Invertebrates Major Characteristics Annelids Mollusks Arthropods Echinoderms Germ Layers Three Three Three Three Body Symmetry Bilateral Bilateral Bilateral Radial (adults) Cephalization Present Present Present Absent (adults) Coelom True coelom True coelom True coelom True coelom Early Development Protostome Protostome Protostome Deuterostome Invertebrate Cladogram Section 29-1 Echinoderms Chordates Arthropods Annelids Mollusks Radial Symmetry Roundworms Flatworms Pseudocoelom Cnidarians Radial Symmetry Protostome Development Three Germ Layers; Bilateral Symmetry Sponges Deuterostome Development Coelom Tissues Multicellularity Single-celled ancestor Pg. 747 Section 29-2 29–2 Section Outline Form and Function in Invertebrates A. Feeding and Digestion 1. Intracellular and Extracellular Digestion 2. Patterns of Extracellular Digestion B. Respiration 1. Aquatic Invertebrates 2. Terrestrial Invertebrates C. Circulation 1. Open Circulatory Systems 2. Closed Circulatory Systems D. Excretion 1. Aquatic Invertebrates 2. Terrestrial Invertebrates E. Response 1. Centralization and Cephalization 2. Specialization F. Movement and Support 1. Hydrostatic Skeletons 2. Exoskeletons 3. Endoskeletons G. Sexual and Asexual Reproduction Section 29-2 Type of Skeleton Types of Invertebrate Skeletons Sponges Hydrostatic X Exoskeleton Endoskeleton Cnidarians Simple (spicules or spongin) Flatworms X Roundworms X Types of Invertebrate Skeletons Section 29-2 continued Type of Skeleton Hydrostatic Exoskeleton Endoskeleton Annelids X Mollusks Arthropods Echinoderms Some Some (shells) X X Section 29-2 Invertebrate Digestive Systems Intestine Gizzard Crop Mouth/anus Pharynx Mouth Gastrovascular cavity Annelid Anus Gastrovascular cavity Cnidarian Pharynx Crop Arthropod Anus Pharynx Mouth Mouth/anus Flatworm Stomach and digestive glands Rectum Intestine Section 29-2 Invertebrate Respiratory Systems Gill Tracheal tubes Siphons Movement of water Insect Mollusk Airflow Book lung Spider Spiracles Section 29-2 Invertebrate Circulatory Systems Heartlike structure Hearts Small vessels in tissues Heart Blood vessels Sinuses and organs Heartlike structures Insect: Open Circulatory System Annelid: Closed Circulatory System Blood vessels Invertebrate Excretory Systems Section 29-2 Flame cells Flatworm Excretory tubules Nephrostome Excretory pore Flame cell Excretory tubule Nephridia Digestive tract Annelid Arthropod Malpighian tubules Section 29-2 Invertebrate Nervous Systems Arthropod Brain Ganglia Ganglia Brain Nerve Cells Flatworm Cnidarian Mollusk Video Circulatory Systems Sponges & Cnidarians Worms & Mollusks Arthropods & Echinoderms Characteristics of Animal Kingdom Heterotrophs Mobile Multicellular Lacks cell walls Sexual reproduction Body symmetry Radial (radiate outward from center) Bilateral (two sides) Dorsal = top side Ventral = belly (bottom) side Anterior = front end Posterior = rear end Asymmetrical (no plane dividing body into equal halves) Which Type of Symmetry? A B B E C F F C D G Development & Body Cavities Zygote (fertilized egg) blastula (hollow ball) gastrula (inward folding creating layers) Ectoderm Endoderm Mesoderm Germ Layers form embryonic layers. Endoderm: inner layer digestive tract and respiratory system Mesoderm: middle layer muscles, circulatory, reproductive and excretory organs Ectoderm: outermost layer sense organs, nerves, and epidermis Body plans Body plans No body cavity Fluid filled body cavity fully lined with mesoderm Fluid filled body cavity partially lined with mesoderm Protostomes vs Deuterostomes Phylum Porifera, the sponges Asymmetrical Lacks tissues = cells embedded in matrix Sessile adults sponges Phylum Cnidaria Examples: jellyfish, coral, hydra, sea anemones, man-o-war Radial symmetry True tissues 2 cell layers endoderm, ectoderm Polyp (tubular body) & Medusa (bell-shaped body) Feeding Body tissue surrounds gastrovascular cavity, a single opening space, where digestion takes place. Phylum Platyhelminthes, the flatworms Examples: planarians, flukes, tapeworms Acoelomates (no cavity) Organs & systems Three germ layers Bilateral symmetry Cephalization (concentration of nervous tissue in head) Flatworms Planeria Intermediate hosts •Tapeworm larvae burrow into cow or fish muscle tissue, which infects the animal that eats this tissue. Rat tapeworm Parasites have a complex life history with several hosts Human tapeworm •Scolex: head with suckers or hooks Parasitic tapeworm from a fish Tapeworm Phylum Nematoda, the roundworms Examples = heartworms, hookworms, pin worms Roundworm Anatomy Pseudocoelom false body cavity between endoderm & mesoderm 2 body openings (mouth and anus) Guinea worm disease Preventive Measures Health education and low-technology measures to promote behavioral change are used to prevent Guinea worm disease. The most effective way to prevent it is to filter the tiny water fleas out of drinking water. The Carter Center provides families with fine-mesh filter cloths that fit over clay pots used to hold water. Some people, especially nomadic groups, receive pipe filters, which are small straw-like personal filters that can be worn around the neck. These simple but revolutionary devices enable people to drink water safely no matter where they are. http://www.cartercenter.org/news/multimedia/media_console/console.aspx?sectionID=H&directoryEXT =h07&linkEXT=CC_video.swf Pinworm infection in child’s intestine Roundworms A report by the Centers for Disease Control and Prevention found that about 14 percent of Americans are infected with Toxocara – a parasite commonly known as the round worm. The disease is zoonotic – which means you can get it from your pet. The CDC says the parasite is most commonly found in children from low socioeconomic areas. For the most part there aren't any symptoms. But in rare cases it can cause blindness, especially in children. Researchers say the worms are transferred through cat and dog fecal matter – in yards and sand boxes – places where children often play. Filarial worms •Primary cause of elephantiasis condition where parts of the body are swollen •Requires arthropod intermediate host to complete life cycle Filarial worms Elephantiasis is a condition of extreme edema that occurs when lymph vessels become blocked by filarial worms. Here the left leg is swollen with the fluids accumulated in the tissues as a result of the blockage. Phylum Annelida Examples: leeches, earthworms & marine worms Segmented worms Separated by septa = dividing walls Bilateral symmetry Coelomates 3 cell layers Advanced organ systems Annelids Leeches release a chemical that prevents the prey from forming scabs. A leech bite is less painful than it is just messy with the profuse bleeding. Phylum Mollusca Coelomates= true body cavity (between mesoderm) Soft body Unsegmented 2nd most successful land animal (after insects) Mollusks Mollusk Anatomy 3 body parts: Visceral mass = houses body organs Mantle = heavy fold of tissue surrounding visceral mass Foot = ventral surface for locomotion Respiratory structure: Gills = filamentous projections that capture oxygen from the water Mollusk Groups Cephalopod = “head-footed” little or no shell most advanced eye Jet propulsion Octopus, squid, nautilus Cuttlefish Mollusk Groups Bivalves = “2 shells” “hatchet-footed” Clams, oysters, etc Mollusk Groups Gastropods = “belly-footed” One shell Snails, slugs, etc Groups of mollusks Gastropods: shell-less or singleshelled mollusks with muscular feet Sea snail Movement Snails: secrete mucus and crawl slowly Cephalopods: fast moving, drawing water into mantle cavity and forcing water out of siphon (like jet propulsion) excessive amount of snail mucus secreted Snail Trail Mexican turbo snail Phylum Arthropoda Jointed appendages Segmented bodies Exoskeleton Largest phylum in Animal Kingdom Metamorphosis Undergoes morphological changes during life history Incomplete = 3 stages Egg, nymph, adult Complete = 4 stages Egg, larva, pupa, adult Arthropod Groups Arachnids = spiders, scorpions Crustaceans = lobsters, shrimp Insects = grasshoppers, butterflies, bees Diplopoda = millipedes Chilopoda = centipedes Specialized appendages Mouthparts Antennae Legs Pincers Wings Swimmerets Phylum Echinodermata Spiny skinned Radial symmetry (as adults) Endoskeleton made of plates, ossicles Water vascular system for moving & feeding Regeneration Water Vascular System, for feeding & locomotion Echinoderm groups Sea Stars (starfish) Sand dollars & sea biscuits Sea urchins Echinoderm groups Sea cucumbers Brittle stars (serpent stars) Sea lilies Echindoerms Subphylum Arachnida – black widow spider Brown Recluse Spider Scorpion Wolf Spider Subphylum Crustacea – hermit crab Subphylum Crustacea –crab Subphylum Crustacea – fire shrimp Subphylum Insecta – lady bug Subphylum Insecta – bee moth The purple Sea urchin has sharp spines to protect against predators. A close-up of the mouth of a Green Sea urchin, called Aristotles lantern. Diversity of Echinoderms Sea cucumbers Sea lilies & feather stars Red Prickly Sea Cucumber Yum Tasty!!!!!!!! Sea Lilies