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Chapter 3 The Diversity of Life Part Two: Classification and Diversity 1 Taxonomic Groups • Six kingdoms – Prokaryotes 1. Archaeobacteria 2. Eubacteria – Eukaryotes 3. Plantae – producer 4. Fungi – consumer 5. Animalia http://www.ucmp.berkeley.edu/ – consumer 6. Protista http://tolweb.org/tree/phylogeny.html 2 Systematics Understanding the history of life Systematics is based on our understanding the life diversified from a single origin. Diversity is a product of descent with modification. • Taxonomy – Naming and classification of life – System of organizing the relationships between organisms • Taxonomic groups – The six kingdoms and their subordinate groups 3 Taxonomic Groups • Taxa range from broad (phylum) to narrow (species) Kingdom Phylum Class Order Family Genus Species Animalia Chordata Mammalia Primates Hominidae Homo Homo sapiens – Species • Group of individuals that can interbreed • Name includes genus • Italicized 4 Taxonomic Groups • Primitive traits – Appear early in evolutionary history – Hagfish group traits • Derived traits – Evolved later – Present only in some subgroups – Jaws, lungs, claws or nails, feather, fur, and mammary glands 5 Prokaryotes- Eubacteria Prokaryotes: Unicellular (mostly) organisms without a nucleus Bacteria may be aerobic or anaerobic Classified as 1. heterotrophs fermentation and respiration 2. autotrophs photoautotrophic or chemoautotrophs • Critical to Earth System as a mechanism for recycling nutrients like carbon, nitrogen and sulfur • Carbon is removed from the atmosphere by plants. Bacteria oxidize this carbon after plants death, which recycles it back to the atmosphere as CO2. No CO2= No plants= No life • Oldest known fossils are 3.5 billion years old: bacteria like organisms • Most common in rock record are stromatolites: Layered mats of cyanobacteria Photo: http://www.sharkbay.org/terrestial_enviroment/page_15.htm 6 Prokaryotes- Archaeans • • Discovered in the 1970’s Inhabit extreme environments – Temperatures >100 degrees Centigrade • Hot springs – Low or no oxygen – Alkaline or Acidic conditions • • They are methanogens: they produce methane Have probably not changed much through time: recognized as far back as 3.8 bya http://www.ucmp.berkeley.edu/archaea/archae asy.html - No driving mechanism for evolution. No competition. 7 Eukaryotes: Protista • Eukaryotes: organisms with complex cells with nuclei. • Protozoans (single celled, nuclei) – Animal-like protists divided on the basis of locomotion and morphology • Amoebas – Change shape • Flagelleates – Flagellum for locomotion • Ciliate – Cilia for locomotion 8 Protista • Unicellular algae – Plant-like protists • Dinoflagellates • Diatoms: chert • Calcareous nannoplankton: chalk • Originated in the Mesozoic Era – Important marine producers 9 Protista Protozoans with skeletons • Foraminifera Chambered skeleton of calcium carbonate Very abundant Useful for dating rocks and sediments Why? They are good index fossils. Planktonic, over 250,000 species, distinct ranges 10 Protista • Protozoa with skeletons • Radiolarians – Skeleton made out of opal (SiO2): chert – Related to foraminifera 11 Eukaryotes: Fungi • Decomposers – Obtain nutrients from dead organisms using digestive enzymes (heterotrophs) – Spore producers • Diverse – Yeast – Mushroom • Poor fossil record 12 Plants • First land plants in the Ordovician (490-443my) • Vascular versus Nonvascular – Vessels for transport of water, dissolved nutrients, food – Transportation of materials by diffusion • Example: Mosses 13 Spore Plants • Ferns – Devonian (417-354my) – Adapted to life on land with roots and leaves – Spores • One set of chromosomes • Fertilized by sperm • Sperm migration requires moisture – Vast Late Paleozoic swamps led to coal formation – Origin of name “Pennsylvanian Period” after coal mines in Pennsylvania 14 Seed Plants • Seeds enabled plants to exploit drier, non coastal habitats • Gymnosperms – Greek word meaning “Naked seed” plants – Conifers • Cone-bearing plants • Eggs are fertilized in cone by pollen – Pollen bears sperm; carried by wind • Dominant in the Mesozoic • Angiosperms – Flowering plants • Pollen carried by pollenators (animals) • Dominant in the Cenozoic http://en.wikipedia.org/wiki/Angiosperm http://www.ucmp.berkeley.edu/seedplants/seedplants.html 15 Animalia • Two groups – Vertebrates • Possess a backbone – Invertebrates • Coelomates – Body cavity housing internal organs 16 Animals- Sponges – Simple invertebrates – Suspension feeds • Strain particles from water • Mostly eat bacteria • Flagella pump water through internal canals – Calcium carbonate or silica spicules support structure • Cambrian - modern 17 Cnidarians • Jellyfish and corals • Polyp and Medusae stage • Radial symmetry and more complex body plan including mouth, tentacles and digestive cavity 18 Arthropods • • Insects, crabs, spiders, lobsters, trilobites Trilobite – Three-lobed body • Central, left-and right- lobed – – – – • • External skeleton Gill-like structure for respiration Legs Primitive eyes Common in Cambrian (560-490 mya) Illustrates that complex life formed early in Earth History and challenges traditional view of evolution as a move from primitive to advanced forms of life. 19 Mollusks • Bivalves (Clams) • Gastropods (snails) • Cephalopods (octopuses) •Shell of aragonite, calcite, or both •All share a common body plan, though they evolved to fit different ecological niches: crawlers, filter feeders, swimmers 20 Mollusks • Cephalopods – Squids, octopuses, chambered nautilusses – Swim in the sea • Jet propulsion • Eyes – Carnivores • Catch with tentacles • Eat with strong beak • Chambered nautilus – Bouyancy due to gas in shell – Pressure accomodated by increased surface area • Common in Phanerozoic 21 Lophophores • Brachiopods – Shell divided into two valves – Lampshells – Lophophores • Pump water • Strain food – Inarticular brachiopods • Lack hinge teeth • Lingula – Articulate brachiopods 22 Bryozoans • Moss animals • Ressemble corals, but more closely related to brachiopods – Lophophore extended from skeleton to feed – Calcified skeleton • Colonial • Ordovician 23 Echinoderms • Spiny-skinned form • Five-fold symmetry – Starfishes • Predators • Lower Paleozoic – Sea urchins • Regular sea urchins – Radially symmetrical bodies • Irregular sea urchins – Bilaterally symmetric – Burrowers 24 Echinoderms • Crinoids – Sea lillies – Sieve food using arms • Pass food to mouth with tube feet – May swim – May be attached by flexible stalk – Disk-shaped plates from stalk 25 Chordates Notochord, Gills and Tail Notochord – Flexible, rodlike structure that supports body for some part of life cycle – Spinal cord • Runs next to notochord 26 Vertebrates 27 Vertebrates • Fishes – Devonian 417 to 354 mya: Age of Fishes – Jawless Fishes “Class Agnatha” – Placoderms: Earliest jawed fishes. Armored plates but no teeth- sharp jaws instead. Devonian – Ray-finned fishes • Fins supported by thin bones radiating from body – Lobe-finned fishes (sharks) • Tetrapods: Amphibians An Ichthyosaur – Amphi meaning both, Bios meaning life – 3000 living species – Devonian: Ichthyostega climbs28to shore in Greenland Vertebrates: Tetrapods • Tetrapods: Reptiles – Eggs with protective shells – Ectothermic • Environment controls internal body temperature • Question: What might be an advantage of this quality? – Dinosaurs • Primitive reptiles 225 to 65 mya – Birds • Endothermic • Evolved from Dinosaurs 29 Vertebrates: Tetrapods • Mammals – Endothermic; Hair – Bear live young – Monotremes • Mammals that lay eggs – Marsupial • Offspring develop in pouch – Placental 30 31 Answers to your questions… • How often do geologists find new species? • What kind of fossils are in N. Idaho? • What is the most common type of fossil found today? • • • • • • • Fungal Spores from the Rhynie Chert, Scotland Devonian age (417-354my) Total named species on Earth: 1.5 million. Actual number may= 100 million (mostly insects), but probably around 13-14 million. (Society for Conservation Biology) Grad student here in department found new species of brachiopod, got to name it. Stephen J. Gould estimated that 99% of all plant and animal species are exinct. Hagerman Fossils (S. Idaho): Pliocene age (3.7-3.1my). 140 species including horses, saber tooth cats, ground sloths… Miocene (20my) Plant Fossils Cambrian Trilobites at Lake Pend Oreille (545-495my) 32