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Transcript
Chapter 22~ • Descent with Modification: A Darwinian View of Life Evolution • Evolution: the change over time of the genetic composition of populations • Natural selection: populations of organisms can change over the generations if individuals having certain heritable traits leave more offspring than others (differential reproductive success) • Evolutionary adaptations: a prevalence of inherited characteristics that enhance organisms’ survival and reproduction November 24, 1859 Triassic Permian 225 Carboniferous 350 Devonian Silurian Ordovician Cambrian Ediacaran 400 430 500 570 700 Precambrian, Proterozoic, & Archarozoic 4500 Life’s Natural History is a record of Successions & Extinctions Seed Plants Land Plants Birds Mammals Reptiles Insects Amphibians Teleost Fish Jawless Fish Chordates Molluscs 280 Flowering Plants 180 Dinosaurs Jurassic 135 Arthropods Cretaceous 63 Multicellular Animals Tertiary Green Algae 1.5 Photosynthetic Bacteria Quaternary Anaerobic Bacteria mya Evolutionary history • • • • • Linnaeus: taxonomy Hutton: gradualism Lamarck: evolution Malthus: populations Cuvier: paleontology • • • • Lyell: uniformitarianism Darwin: evolution Mendel: inheritance Wallace: evolution Charles Darwin • 1809-1882 • British naturalist • Proposed the idea of evolution by natural selection • Collected clear evidence to support his ideas Voyage of the HMS Beagle • Invited to travel around the world – 1831-1836 (22 years old!) – makes many observations of nature • main mission of the Beagle was to chart South American coastline Stopped in Galapagos Islands 500 miles off coast of Ecuador Darwin found… birds Collected many different birds on the Galapagos Islands. Thought he found Finch? very different kinds… Sparrow? QuickTime™ and a Photo - JPEG decompressor are needed to see this picture. Woodpecker? Warbler? But Darwin found… a lot of finches Darwin was amazed to find out: All 14 species of birds were finches… But there is only one species of finch on the mainland! Finch? Sparrow? Large Ground Small Ground Finch? Sparrow? QuickTime™ and a Photo - JPEG decompressor Finch Finch are needed to see this picture. How did one species of finches become so many different species now? Woodpecker? Warbler Finch Woodpecker? Warbler? Veg. Tree Finch Warbler? Darwin’s finches • Differences in beaks – associated with eating different foods – survival & reproduction of beneficial adaptations to foods available on islands Warbler finch Cactus finch Woodpecker finch Sharp-beaked finch Small insectivorous tree finch Large insectivorous tree finch Small ground finch Cactus eater Medium ground finch Insect eaters Seed eaters Vegetarian tree finch Bud eater Large ground finch Darwin’s finches • Darwin’s conclusions – small populations of original South American finches landed on islands • variation in beaks enabled individuals to gather food successfully in the different environments – over many generations, the populations of finches changed anatomically & behaviorally • accumulation of advantageous traits in population • emergence of different species Seeing this gradation & diversity of structure in one small, intimately related group of birds, one might really fancy that from an original paucity of birds in this archipelago, one species has been taken & modified for different ends. Darwin’s finches • Differences in beaks allowed some finches to… – successfully compete – successfully feed – successfully reproduce • pass successful traits onto their offspring More observations… Correlation of species to food source Whoa, Turtles, too! Essence of Darwin’s ideas • Natural selection – variation exists in populations – over-production of offspring • more offspring than the environment can support – competition • for food, mates, nesting sites, escape predators – differential survival • successful traits = adaptations – differential reproduction • adaptations become more common in population Evolution evidence: Biogeography • Geographical distribution of species • Examples: Islands vs. Mainland Australia Continents Evolution evidence: The Fossil Record • Succession of forms over time • Transitional links • Vertebrate descent Fossil Record 2006 Fossil Discovery of Early Tetrapod • Tiktaalik – “missing link” from sea to land animals Evolution evidence: Comparative Anatomy • Homologous structures (homology) • Descent from a common ancestor • • • • Homologous structures Similar structure Similar development Different functions Evidence of close evolutionary relationship – recent common ancestor Homologous structures spines leaves succulent leaves needles colored leaves tendrils Analogous structures Separate evolution of structures similar functions similar external form different internal structure & development different origin no evolutionary relationship Don’t be fooled by their looks! Solving a similar problem with a similar solution Vestigial organs • Modern animals may have structures that serve little or no function – remnants of structures that were functional in ancestral species – deleterious mutations accumulate in genes for noncritical structures without reducing fitness • snakes & whales — remains of pelvis & leg bones of walking ancestors • eyes on blind cave fish • human tail bone Evolution evidence: Comparative Embryology • Pharyngeal pouches, ‘tails’ as embryos Evolution evidence: Molecular Biology • Similarities in DNA, proteins, genes, and gene products • Common genetic code Closely related species have sequences that are more similar than distantly related species DNA & proteins are a molecular record of evolutionary relationships Building “family” trees Closely related species (branches) share same line of descent until their divergence from a common ancestor Artificial selection • Artificial breeding can use variations in populations to create vastly different “breeds” & “varieties” “descendants” of wild mustard “descendants” of the wolf Natural selection in action • Insecticide & drug resistance – insecticide didn’t kill all individuals – resistant survivors reproduce – resistance is inherited – insecticide becomes less & less effective Final words…... • “Absence of evidence is not evidence of absence.”