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Ecology Campbell, Chapters 50-54 Ecology v. Environment • Hierarchy of Ecology – biosphere – ecosystem – community – population – organismal Biotic and Abiotic Factors • components of the environment • biotic – other species may affect one species and its distribution • abiotic – temperature – water – sunlight – Wind – rocks and soil Population Ecology • Density – mark and recapture method – line intercept method – point count method • Distribution – clumped – uniform – random Age Distribution • distribution of males and females in each age group of a population • used to predict future population growth Survivorship • mirrors mortality • expressed in survivorship curves • plots surviving individuals at different age groups • three types of survivorship curves • late loss (Type I) • constant loss (Type II) • early loss (Type III) Population Dynamics: Exponential Model • population lives in an area with no environmental limitations N B D t B = number of births in time period D = number of deaths in time period • example – # of births over one year is 500 and # of deaths over one year is 50 N 500 50 450 individuals t Population Dynamics: Exponential Model • per capita rate of increase – average number of births and deaths calculated as rates • J curve • exponential growth r bd # of births b N # of deaths d N Population Dynamics: Exponential Model • example – 10,000 birds in a population – 1500 births and 500 deaths per year – 1500/10,000 - 500/10,000 = .10 or 10% – expressed by stating there is a 10% increase per bird per year Population Dynamics: Exponential Model • measures optimal population growth • rmax = intrinsic rate of increase rmax N t • example – if N=500; r=.05 Calculate the intrinsic rate of increase N .05500 25 individuals t Population Dynamics: Logistic Growth Model • size of a population is limited to: – intrinsic rate of increase – environmental resistance • includes limitations the environment imposes on birth rate and death rate in a population – – – – food space predation Parasitism • carrying capacity (K) determined by – renewable resources like water, nutrients, and light – nonrenewable resources such as space Population Dynamics: Logistic Growth Model – logistic population growth – r decreases as N increases – K-N tells us # of individuals population can accommodate – S curve N KN rmax ( )N t K Logistic Growth Model and Life Histories • K-selected – equilibrial populations – live at density near limit imposed by resources • r-selected – opportunistic populations – live in environments where little competition is present Density Influence on Birth and Death Rates • density independent – unrelated to population size – most important are • weather • Climate • density dependent – increase effectiveness as population density increases – especially affects long lived organisms – include • predation • parasitism • competition Human Population Growth • J curve growth • grows at a rate of about 80 million yearly (r=1.3%) • Why doesn’t environmental resistance take effect? • altering their environment • technological advances – the cultural revolution – the agricultural revolution – the industrial-medical revolution Community Ecology • competition • predator-prey • symbiosis – mutualism – commensalism – parasitism – amensalism • species richness • relative abundance • species diversity Competition • intraspecific competition • interspecific competition • Competitive exclusion principle Predator-prey Interactions • one species eating another – herbivory – carnivory – parasitism – cannibalism • predators are agents of natural selection • animal defenses against predators – hiding or escape – mimicry – adaptive coloration Mimicry • Batesian mimicry Mimicry • Müllerian mimicry Mimicry • aggressive mimicry Adaptive Coloration • cryptic coloration Adaptive Coloration • deceptive coloration Adaptive Coloration • aposematic coloration Symbiosis • involves – host – symbiont • 3 divisions – mutualism – commensalism – parasitism Mutualism - lichens Symbiosis • involves – host – symbiont • 3 divisions – mutualism – commensalism – parasitism Commensalism – sea slug and chloroplasts Symbiosis • involves – host – symbiont • 3 divisions – mutualism – commensalism – parasitism Parasitism – Chagas’ Disease Coevolution • reciprocal evolutionary change – e.g. flower shape and pollination • creates a “balance of nature” Moderator of Competition • keystone species – removal dramatically alters the environment – influences species diversity Trophic Structure • feeding relationships • trophic levels – primary producers – primary consumers (herbivores) – secondary consumers (primary carnivores) – tertiary consumers (secondary carnivores) • detritovores • omnivores • food chain vs. food web Food Chain v. Food Web Ecological Succession • structural change in a community and its nonliving environment over time • typically occurs as a result of some environmental disturbance • 2 forms – primary succession • begins in a lifeless area with invaders called pioneers • may proceed to a stable climax community • vary according to geography, climate, etc – secondary succession • occurs in an existing community where a disturbance has occurred Ecosystem Processes • Production • Consumption • Decomposition Trophic Relationships exhibit all 3 processes – autotrophs produce – heterotrophs consume – detritovores and decomposers recycle and decompose Production: Energy Flow • light energy – about 1% of sun’s energy used for life – converted via photosynthesis • How do we measure the amount of energy incorporated by producers? – primary production Consumption: Energy Flow • ecological efficiency – 10% on average • energy pyramid • biomass pyramid Nutrient Cycles • Biogeochemical cycles – materials move through biotic and abiotic portions – reservoir Biogeochemical Cycles • Be able to answer the following questions about each biogeochemical cycle • 1. What is the reservoir? • 2. How does the nutrient enter the biotic portion of the cycle? • 3. How is the nutrient returned to the reservoir? Biological Magnification • toxic substances accumulate in higher concentrations as trophic levels increase • toxin is usually not biodegradable • result of inefficiency of energy transfer Human Impact: Deforestation Human Impact: Global Warming Human Impact: Global Warming Human Impact: Acid Rain Human Impact: Agriculture • manipulate ecosystem productivity – use of herbicides to get rid of unwanted plants – use of pesticides to get rid of unwanted pests – fertilization and irrigation – genetic modification of plants to increase yield