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The Theory of Evolution • • • • • Charles Darwin Evidence for evolution Mechanisms for evolution Natural selection Speciation Evidence of evolution • Structural adaptations – Mimicry – Camouflage • Physiological adaptations – Change in a metabolic process • Fossils • Anatomy – Homologous structures – Analogous structures – Vestigial structure – Embryos • Biochemistry http://evolution.berkeley.edu/evolibrary/home.php Theory of Natural Selection The Theory of Evolution • • • • • • Observation: Charles Darwin Evidence for evolution Mechanisms for evolution Natural selection Speciation Limits are placed on population expansion by limited environmental resources • Conclusion: Therefore not all organisms will survive to adulthood and reproduce – therefore there must be a “struggle for existence” (Source: observations reinforced by Malthus) http://evolution.berkeley.edu/evolibrary/home.php 1 Theory of Natural Selection Theory of Natural Selection Conclusions: Observations: • Not all members of a species are alike (variation) • Parents often pass their individual variations on to their offspring • individuals featuring favorable variations will enjoy a competitive advantage over others • they will survive in proportionately greater numbers and will produce offspring in increasingly greater numbers (Source: Animal breeders, taxonomists) Theory of Natural Selection There is “differential reproduction” and “differential survival,” i.e., “natural selection” (Author of inference: Darwin) Theory of Natural Selection Conclusion: • Through the action of natural selection over many generations a species could evolve (Author of inference: Darwin) 2 Theory of Natural Selection Both Darwin and Wallace knew: the principle cause of natural selection is the environment Biston betularia Theory of Natural Selection (a) The dark form is more visible on the light, lichencovered tree. (b) On trees darkened by pollution, the lighter form is more visible. Variation in the peppered moth. Understanding Physical Anthropology and Archaeology, 9th ed., p. 29 But look at them without color vision. 3 Theory of Natural Selection • The moths satisfy all four conditions for natural selection: • they reproduce; • their color pattern is inherited; • there is variation in their color patterns; • the different forms have different fitness. Mechanisms for evolution • Do populations or individuals evolve? Populations • What is a gene pool? All of the alleles in a population • How can the gene pool change? – Mutation – Genetic drift – Gene flow • Would these things effect a large population or a small population more? Contrasting ideas about the mechanism of evolution. (a)According to Lamarck’s theory (b) According to the Darwin-Wallace Understanding Physical Anthropology and Archaeology, 9th ed., p. 24 4 The Theory of Evolution • • • • • Charles Darwin Evidence for evolution Mechanisms for evolution Natural selection Speciation http://evolution.berkeley.edu/evolibrary/home.php What is speciation? What is speciation? • Geographic isolation • Reproductive isolation • • • • Geographic isolation Reproductive isolation Gradualism Punctuated equilibrium http://evolution.berkeley.edu/evolibrary/home.php 5 What is speciation? • • • • • Geographic isolation Reproductive isolation Gradualism Punctuated equilibrium Divergent evolution Adaptive radiation – an example of divergent evolution – Adaptive radiation • Convergent evolution http://www.vanderbilt.edu/AnS/english/Clayton/G alapago_finches.gif http://evolution.berkeley.edu/evolibrary/home.php Convergent evolution – what is an example? Theory of Natural Selection • Problem What is the source of individual variation? lack of knowledge of modern genetics http://evolution.berkeley.edu/evolibrary/home.php 6 Theory of Natural Selection Neither Darwin nor Wallace knew: the source of individual variation = genetics (inherited characteristics) Theory of Natural Selection Problem: If natural selection only weeds out what already exists, how can it produce anything new? Didn’t know then because of lack of knowledge of mutation and sexual recombination from modern genetics Mechanisms for evolution • Do populations or individuals evolve? • What is a gene pool? • How can the gene pool change? – Mutation – Genetic drift – Gene flow • Would these things effect a large population or a small population more? 7