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THEORY OF EVOLUTION Chapter 15 15-1 History of Evolutionary Thought I. Charles Darwin (1809 – 1882) A. Studied to be a doctor, then a minister, but was also a naturalist B. In 1830’s traveled around world for five years on the HMS Beagle C. Went to Galapagos Islands and South America D. Noticed that Galapagos tortoises looked slightly different on each island E. Also noted that plant/animal species resembled those on South American mainland but were not the same F. Gathered & recorded lots of data (plants, animals, fossils) G. Formulated idea of natural selection to explain how species evolve, or change over time 1. Used his data and the ideas of other scientists H. Alfred Wallace – Darwin’s contemporary also had same ideas, but Darwin published his manuscript first, On the Origin of Species by Means of Natural selection(1859) II. Ideas of Darwin’s Time A. In 1700’s, most scientists thought species were fixed and unchanging and that Earth was 1000’s of years old, not billions. Not to mention that it was considered heresy to go against the teachings of the church (religion). B. Nicolaus Steno identified rock strata (layers) and that lower layers were older “Law of Superposition” 1. layers had fossils of different types of organisms C. Georges Cuvier – reconstructed fossils and proposed that some species were extinct 1. deeper strata had fossils that were really different from modern orgs 2. found “sudden” changes in kinds of organisms found in rock strata 3. proposed catastrophism – sudden geologic catastrophes caused extinction of species 4. Not all of Cuvier’s ideas are accepted, but geologic change & extinction are! D. Charles Lyell – proposed uniformitarianism – same geological processes that shaped Earth in the past are still at work now E. Jean Baptiste Lamarck 1. simple organisms could come from nonliving things (not true) 2. simpler organisms develop into more complex ones (not true) 3. inheritance of acquired characteristics – traits acquired during one’s life are passed down (not true) Lamarck was important even though his ideas were wrong, he was the first to propose evolution but his proposed mechanisms for evolution in organisms were wrong 15-2 Evidence of Evolution I. Fossil Record • Provides evidence of Evolution • Fossil Record – fossils found in various layers of strata. Some fossils have a wide geographic distribution within a specific time. These fossils are used to estimate the time of other less common fossils found in the same strata (relative age) • Fossils are remains of once living organisms and are found only in sedimentary rock Cast Fossil Remineralized fossil Amber Fossil A. Age of Fossils • Nicolaus Steno (1669) – Principle of Superposition – lower strata have fossils of older organisms than higher strata . • Oldest layers on bottom, youngest layers on top. Relative dating v. Absolute dating – relative dating estimates the age of a fossil based on the layer of the stata that the fossil is found in and other fossils found in that strata – Absolute dating – is done by radiometric dating, this provides a much more accurate age of the fossil or material. C. Transitional Species • Fossils show that there were species that were intermediate between those of their ancestors and descendants • Can show the sequence of evolution of different characteristics • Example – evolution of whales from four-legged, land mammals (p. 304) II. Biogeography • Biogeography is the study of the location and distribution of organisms on Earth. • Closely related species tend to live near each other on Earth & distantly related species tend to live far apart Example: • Australian sugar-glider looks like a North American flying squirrel, but is more closely related to other marsupials on Australia • they have similar adaptations to their similar environments (convergent evolution ) • Sugar-glider is a marsupial because it’s ancestors were marsupials • Convergent evolution – two unrelated species have similar adaptations because they are exposed to same type of environment. III. Comparative Anatomy and Embryology • Comparitive Anatomy: • Homologous structures - show common ancestry (similar form, not necessarily same function). So the underlying morphology is the same (look at the bones and structure) • Example: forelimb bones of human, bat, whale Analogous structures • - DO NOT show common ancestry, but have similar function Vestigial structures • Have no function in a species, but resemble structures that do have function in related species or that did have a function in ancestors Example: some snakes and whales have vestigial remains of pelvis & leg bones that were present in their walking ancestors Example: human tailbone resembles those bones in animals with tails B. Comparative Embryology • Embryology – the study of how an organism develops • Similar species resemble each other as they develop and decrease their resemblance as development proceeds IV. Molecular Evidence • Species with more similar DNA, RNA, and/or proteins are more closely related than species with fewer DNA sequences, and proteins in common. 15-3 Evolution in Action I. Divergence and Radiation • Divergent evolution – many species evolve from a common ancestor as they spread (radiate) out and occupy different environments or ecological niches (niche – everything about an organisms life, what it eats, where it lives, and how it finds a mate) Example: adaptive radiation – finches or tortoises on Galapagos . • Contrast convergent evolution with adaptive radiation – • Convergent evolution - – unrelated species evolve similar traits because they live in similar environments • Adaptive radiation – one species spreading out to occupy different ecological niches to deal with competition within the species. II. Evolution in Action • Co-evolution – two species evolve adaptations as a result of each other’s influence. Think two species that rely on one another. Predator and prey Flowers and polinators • Artificial Selection – this is a process done by human control over which individuals breed and produce offspring based on desired traits. • Sexual Selection – some individuals have better success at obtaining a mate. So they also have better reproductive success. • Usually the female of the species drives sexual selection.