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11.1 Genetic Variation Within Population KEY CONCEPT A population shares a common gene pool. 11.1 Genetic Variation Within Population SPECIES- group of organisms so similar that they can breed to produce fertile offspring POPULATION- a group of the same species living in an area where no two individuals are exactly alike due to variations that have led the fittest individuals to survive and pass on these traits ALLELE- An allele is an alternative form of a gene. Organisms typically have two alleles for a single trait, one being inherited from each parent. ALLELE FREQUENCY- measure of how common an allele is in a population GENE POOL- all of the alleles of all species in a population 11.1 Genetic Variation Within Population Genetic variation in a population increases the chance that some individuals will survive. • Phenotype-a trait produced by 1 or more genes which provides variation in the gene pool. Examples: body size- tall or thin; feather patterns- ruffled or smooth • Genetic variation leads to phenotype variation. • Phenotype variation is necessary for natural selection. • Genetic variation is stored in a population’s gene pool. – Gene pool is made up of all alleles in a population – New allele combinations form when organisms have offspring 11.1 Genetic Variation Within Population • Allele frequencies measure genetic variation. – measures how common allele is in population – can be calculated for each allele in gene pool 11.1 Genetic Variation Within Population Genetic variation comes from several sources. • Mutation is a random change in the DNA of a gene. – can form new allele – can be passed on to offspring if in reproductive cells • Recombination forms new combinations of alleles. – usually occurs during meiosis – parents’ alleles arranged in new ways in gametes 11.1 Genetic Variation Within Population Genetic variation comes from several sources. • Hybridization is the crossing of two different species. – occurs when individuals can’t find mate of own species – topic of current scientific research 11.1 Genetic Variation Within Population KEY CONCEPT Populations, not individuals, evolve. 11.1 Genetic Variation Within Population Natural selection acts on distributions of traits. • A normal distribution graphs as a bell-shaped curve. – highest frequency near mean value (MEAN VALUE IS THE AVERAGE VALUE) – frequencies decrease toward each extreme value SO…EXTREME VALUES LIE TOWARD THE BOTTOM OF EACH SIDE OF THE TOP• Traits not undergoing MIDDLE MEAN/AVERAGE natural selection have a VALUES normal distribution. 11.1 Genetic Variation Within Population • Microevolution is evolution within a population from generation to generation – observable change in the allele frequencies – can result from natural selection 11.1 Genetic Variation Within Population Natural selection can change the distribution of a trait in one of three ways. 1) Directional Selection 2) Stabilizing Selection 3) Disruptive Selection These types of selection shifts cause a certain phenotype trait to become more common in the population 11.1 Genetic Variation Within Population 1) Directional selection favors phenotypes at one extreme of a trait’s range causing it to shift in that direction. 11.1 Genetic Variation Within Population 2) Stabilizing selection favors the intermediate phenotype leading to a decline of extreme phenotypes and possibly extinction. 11.1 Genetic Variation Within Population 3) Disruptive selection favors both extreme phenotypes. • Favoring both extreme phenotypes can lead to development of new species thru speciation 11.1 Genetic Variation Within Population KEY CONCEPT Natural selection is not the only mechanism through which populations evolve. 11.1 Genetic Variation Within Population Gene flow is the movement of alleles between populations. • Gene flow occurs when individuals join new populations and reproduce. • Gene flow keeps neighboring populations similar. • Low gene flow increases the chance that two populations will evolve into different species. bald eagle migration 11.1 Genetic Variation Within Population Genetic drift is a change in allele frequencies due to chance. • Genetic drift causes a loss of genetic diversity. • It is most common in small populations. • A population bottleneck can lead to genetic drift. – It occurs when an event drastically reduces population size. – The bottleneck effect is genetic drift that occurs after a bottleneck event. – Leads to lack of genetic genetic variation b/c of the minimized population 11.1 Genetic Variation Within Population • The founding of a small population can lead to genetic drift. – It occurs when a few individuals start a new population. – The founder effect is genetic drift that occurs after start of new population. 11.1 Genetic Variation Within Population • Genetic drift has negative effects on a population. – less likely to have some individuals that can adapt – harmful alleles can become more common due to chance 11.1 Genetic Variation Within Population Sexual selection occurs when certain traits increase mating success. • Sexual selection occurs due to higher cost of reproduction for females. – males produce many sperm continuously – females are more limited in potential offspring each cycle b/c they produce only 1 egg 11.1 Genetic Variation Within Population • There are two types of sexual selection. – intrasexual selection: competition among males – intersexual selection: males display certain traits to females 11.1 Genetic Variation Within Population KEY CONCEPT Hardy-Weinberg equilibrium provides a framework for understanding how populations evolve. 11.1 Genetic Variation Within Population Hardy-Weinberg equilibrium describes populations that are not evolving. • Biologists use models to study populations. • Hardy-Weinberg equilibrium is a type of model. 11.1 Genetic Variation Within Population Hardy-Weinberg equilibrium describes populations that are not evolving. • Genotype frequencies stay the same if five conditions are met. – very large population: no genetic drift – no emigration or immigration: no gene flow – no mutations: no new alleles added to gene pool – random mating: no sexual selection – no natural selection: all traits aid equally in survival 11.1 Genetic Variation Within Population Hardy-Weinberg equilibrium describes populations that are not evolving. • Real populations rarely meet all five conditions. – Real population data is compared to a model. – Models are used to studying how populations evolve. 11.1 Genetic Variation Within Population The Hardy-Weinberg equation is used to predict genotype frequencies in a population. • Predicted genotype frequencies are compared with actual frequencies. – used for traits in simple dominant-recessive systems – must know frequency of recessive homozygotes – p2 + 2pq + q2 = 1 "The Hardy-Weinberg equation is based on Mendelian genetics. It is derived from a simple Punnett square in which p is the frequency of the dominant allele and q is the frequency of the recessive allele." 11.1 Genetic Variation Within Population There are five factors that can lead to evolution. 1)Genetic Drift a. Bottleneck effect b. Founders effect 2)Gene Flow 3)Mutations 4)Sexual Selection 5)Natural Selection 11.1 Genetic Variation Within Population • Genetic drift changes allele frequencies due to chance alone via the bottleneck effect and founders effect. 11.1 Genetic Variation Within Population • Gene flow moves alleles from one population to another. • Gene flow is the migration of alleles between populations. 11.1 Genetic Variation Within Population • Mutations produce the genetic variation needed for evolution. 11.1 Genetic Variation Within Population • Sexual selection selects for traits that improve mating success. 11.1 Genetic Variation Within Population • Natural selection selects for traits advantageous for survival. 11.1 Genetic Variation Within Population • In nature, populations evolve. – expected in all populations most of the time – respond to changing environments 11.1 Genetic Variation Within Population KEY CONCEPT New species can arise when populations are isolated. 11.1 Genetic Variation Within Population The isolation of populations can lead to speciation. • SPECIATION: evolution of two or more species from one species • Populations become isolated when there is no gene flow. – Isolated populations adapt to their own environments. – Genetic differences can add up over generations leading to the development of new species 11.1 Genetic Variation Within Population SPECIATION IN CATS 11.1 Genetic Variation Within Population SPECIATION OF GALAPAGOS FINCHES 11.1 Genetic Variation Within Population Types of Isolation that lead to speciation: • A) REPRODUCTIVE ISOLATION • B) BEHAVIORAL ISOLATION • C) GEOGRAPHICAL ISOLATION • D) TEMPORAL ISOLATION 11.1 Genetic Variation Within Population • Reproductive isolation can occur between isolated populations. – members of different populations cannot mate successfully – final step to becoming separate species is the inability to reproduce fertile offspring • Speciation is the rise of two or more species from one existing species. 11.1 Genetic Variation Within Population Populations can become isolated in several ways. • Behavioral barriers can cause isolation. – called behavioral isolation – includes differences in courtship or mating behaviors 11.1 Genetic Variation Within Population • Geographic barriers can cause isolation. – called geographic isolation – physical barriers divide population – Lakes, mountains, roads, canals 11.1 Genetic Variation Within Population Temporal barriers can cause isolation called temporal isolation • Timing of reproductive periods prevents mating • These differences can be time of day, season, or different years, feeding times, etc 11.1 Genetic Variation Within Population Populations can become isolated in several ways. 11.6 Patterns in Evolution KEY CONCEPT Evolution occurs in patterns. 11.6 Patterns in Evolution Evolution through natural selection is not random. • Natural selection can have direction. • The effects of natural selection add up over time. 11.6 Patterns in Evolution • This happens because they face similar environmental conditions or pressure 11.6 Patterns in Evolution 11.6 Patterns in Evolution • Divergent evolution describes evolution toward different traits in closely related species. kit fox red fox ancestor 11.6 Patterns in Evolution 11.6 Patterns in Evolution Species can shape each other over time. • Two or more species can evolve together through coevolution. – evolutionary paths become connected – species evolve in response to changes in each other 11.6 Patterns in Evolution • Coevolution can occur in beneficial relationships. 11.6 Patterns in Evolution • Coevolution can occur in competitive relationships, sometimes called evolutionary. 11.6 Patterns in Evolution Species can become extinct. • Extinction is the elimination of a species from Earth. • Background extinctions occur continuously at a very low rate. – occur at roughly the same rate as speciation – usually affects a few species in a small area – caused by local changes in environment & man-made causes 11.6 Patterns in Evolution 11.6 Patterns in Evolution • Mass extinctions are rare but much more intense. – destroy many species at global level – thought to be caused by catastrophic events – at least five mass extinctions in last 600 million years 11.6 Patterns in Evolution Speciation often occurs in patterns. • A pattern of punctuated equilibrium exists in the fossil record. – theory proposed by Eldredge and Gould in 1972 – episodes of speciation occur suddenly in geologic time – followed by long periods of little evolutionary change – revised Darwin’s idea that species arose through gradual transformations 11.6 Patterns in Evolution • Many species evolve from one species during adaptive radiation. – ancestral species diversifies into many descendent species – descendent species usually adapted to wide range of environments 11.6 Patterns in Evolution 11.6 Patterns in Evolution